Prof. Dr. helene richter
Tel: (+49)-(0)251-83-21014
Fax: (+49)-(0)251-83-23896
Room 154
Office hours: upon consultation
Research Foci
- Refinement of animal experiments and housing conditions
- Reproducibility and generalizability of animal experiments
- Animal welfare
- Cognition and emotion
- Animal welfare and animal personality
CV
Academic Education
- PhD-studies, Animal Welfare and Ethology, Justus-Liebig-Universität Gießen
- Studies of Biology, Westfälische Wilhelms-Universität Münster
Positions
- Professor for Behavioural Biology and Animal Welfare
- Postdoctoral researcher, Department of Behavioural Biology, Westfälische Wilhelms-Universität Münster
- Postdoctoral researcher, Animal Models in Psychiatry, Central Institute of Mental Health and Bernstein Center for Computational Neuroscience (BCCN), Mannheim / Heidelberg
- Postdoctoral researcher, Animal Welfare and Ethology, Veterinary Medicine, Justus-Liebig-Universität Gießen
- PhD student, Animal Welfare and Ethology, Veterinary Medicine, Justus-Liebig-Universität Gießen
Honors
- Margarete von Wrangell-Habilitationsprogramm – Ministerium für Wissenschaft, Forschung und Kunst Baden-Württemberg
- KlarText – Klaus Tschira Preis für verständliche Wissenschaft in the category Fach Biologie (1st place) – Klaus Tschira Stiftung gGmbH
External Functions
- University of Münster (Tierschutzausschuss, Vice speaker)
- Faculty of Biology, University of Münster (Gleichstellungskommission, Speaker)
- Westfälische Wilhelms-Universität Münster (Center for Philosophy of Science, Board member)
- CRC TRR 212 “A Novel Synthesis of Individualisation across Behaviour, Ecology and Evolution: Niche Choice, Niche Conformance, Niche Construction (NC3) (Steering Committee, Member)
- University of Münster (Otto Creutzfeldt Center for Cognitive and Behavioral Neuroscience, Board member)
- Research Training Group EvoPAD ("Evolutionary Processes in Adaptation and Disease") (Steering Committee, Member)
- Manager of the Ethological Society e.V. (Geschäftsführerin)
- Executive board member of the Otto Creutzfeldt Center for Cognitive and Behavioral Neuroscience (OCC), Westfälische Wilhelms-Universität, Münster (Vice speaker)
- Member of the Coordination Commission for Animal Experimentation, University of Münster
- Member of the Hessian Animal Welfare Committee
- Member of the Human Affectome Project, Neuroqualia, Happiness Team, Nova Scotia, Canada
- Member of the Young North Rhine-Westphalian Academy of Sciences, Humanities and the Arts (“Junges Kolleg” NRW)
- Member of the Ethological Society e.V.
- Member of International Society of Applied Ethology (ISAE)
Appointments
- Ruf auf eine Professur für "In vitro und in vivo Modelle" (§ 98 UG 2002), Veterinärmedizinische Universität Wien
University of Veterinary Medicine Vienna, In vitro und in vivo Modelle – rejected - Ruf auf eine Professur für Verhaltensbiologie und Tierschutz (W2), Westfälische Wilhelms-Universität Münster
University of Münster, Verhaltensbiologie und Tierschutz (W2) – accepted - Call for a temporary W2-S-professorship for “Animal welfare and refinement in animal experimentation”, Freie Universität Berlin and German Federal Institute for Risk Assessment (BfR)
Freie Universität Berlin, Animal welfare and refinement in animal experimentation – rejected - Call for a W1-professorship for “Ethology of Farm Animals”, Landwirtschaftliche Fakultät, Rheinische-Friedrich-Wilhelms Universität Bonn
University of Bonn, Ethology of Farm Animals (W1) – rejected
Projects
In Process
- Animal Personality Meets Personality Psychology ()
Internally at the University of Münster Funded Project: | Uni Münster-internal funding - Topical Programmes - Creating an interdisciplinary publication platform for international replication research ()
Internally at the University of Münster Funded Project: | Uni Münster-internal funding - Topical Programmes
Finished
- Rechts oder links? Emotionen, Lateralität und Persönlichkeit bei Hunden – WAU II ()
Individual Granted Project: | Josefine-Charlotte-Schiffer-Stiftung - Individualisierung in sich ändernden Umwelten – InChangE ()
Participation in other joint Project: | MKW - Förderlinie "Profilbildung" | Project Number: PROFILNRW-2020-143-B - 3T: Animal Protection, Animal Welfare, Animal Ethics ()
Internally at the University of Münster Funded Project: | Uni Münster-internal funding - Topical Programmes - The Individual in the Focus of Life Sciences ()
Internally at the University of Münster Funded Project: | Uni Münster-internal funding - Topical Programmes - RTG 2220: Evolutionary Processes in Adaptation and Disease – RTG 2220 EvoPAD ()
Main DFG-Project Hosted at the University of Münster: | DFG - Research Training Group | Project Number: GRK 2220/1 - CRC TRR - 212 A02: Optimistic and pessimistic decision making under ambiguity: causes and consequences for niche choice and niche conformance ()
Subproject in DFG-Joint Project Hosted outside the University of Münster: | DFG - Collaborative Research Centre | Project Number: TRR 212/1 A02 - Weidebilder - Kunst in der Ambivalenz der Mensch-Tier-Beziehung ()
Individual Granted Project: | Bezirksregierung Münster | Project Number: 48.07.16-15/20 RKP - CRC TRR 58 - A01: Life history, serotonin transporter genotype and anxiety-like behaviour: testing the "coping-with-challenge hypothesis" ()
Subproject in DFG-Joint Project Hosted at the University of Münster: | DFG - Collaborative Research Centre | Project Number: TRR 58/4 - Standardisierung, Heterogenisierung und Reproduzierbarkeit von Ergebnissen aus Tierversuchen ()
Individual Granted Project: | DFG - Individual Grants Programme | Project Number: RI 2488/3-1
- Animal Personality Meets Personality Psychology ()
Publications
- Quante, SM, Groneberg, E, Kaiser, S, and Richter, SH. . “The sound of silence: sex and individual differences in rat ultrasonic vocalization.” Behavioural Brain Research 496 115823. doi: 10.1016/j.bbr.2025.115823.
- Mundinger, C, Schulz, NKE, Singh, P, Janz, S, Schurig, M, Seidemann, J, Kurtz, J, Müller, C, Schielzeth, H, von Kortzfleisch, VT, and Richter, SH. . “Testing the reproducibility of ecological studies on insect behavior in a multi-laboratory setting identifies opportunities for improving experimental rigour.” PLoS Biology 23 (4) e3003019. doi: 10.1371/journal.pbio.3003019.
- Carranza, Pinedo V, Krohs, U, and Richter, SH. . “Towards a scientific definition of animal emotions: Integrating innate, appraisal, and network mechanisms.” Neuroscience and Biobehavioral Reviews 172 106127. doi: 10.1016/j.neubiorev.2025.106127.
- Richter, SH, Caspers, B, Dammhahn, M, and Kaiser, S. . “Animal research revisited - the case of behavioural studies.” Trends in Ecology & Evolution 40 (2): 99–103. doi: 10.1016/j.tree.2024.11.014.
- Bračić, M, Bierbaum, L, Peng, M, Nimalavachchlan, L, Siewert, V, Kaiser, S, Sachser, N, and Richter, SH. . “The behavioural ecology of optimism: judgement bias and foraging under predation risk in mice.” Animal Behaviour 219 122991. doi: 10.1016/j.anbehav.2024.10.005.
- Siewert, V, Kaiser, S, Sachser, N, and Richter, SH. . “Optimism and pessimism: a concept for behavioural ecology.” Biological Reviews of the Cambridge Philosophical Society 100: 1152–1162. doi: 10.1111/brv.13178.
- Groneberg, E, Richter, SH, and Kaiser, S. . “Task complexity matters: investigating behavioural lateralization in rats.” Animal Behaviour 123118. doi: 10.1016/j.anbehav.2025.123118.
- Quante, SM, Mundinger, C, Palme, R, Kaiser, S, and Richter, SH. . “What it is like to be an optimist: temporal stability of cognitive bias in rats and its link to other individual traits.” Animal Behaviour 224 123202. doi: 10.1016/j.anbehav.2025.123202.
- Groneberg, E, Richter, SH, and Kaiser, S. . “Is Paw Preference in Rats Influenced by Environmental Novelty?” Ethology 131: 77–93. doi: 10.1111/eth.13580.
- SH, Richter. . “Challenging current scientific practice: How a shift in research methodology could reduce animal use.” Lab Animal 53: 9–12. doi: 10.1038/s41684-023-01308-9.
- Rystrom, TL, Wesseler, Y, Richter, SH, Sachser, N, and Kaiser, S. . “Shaped by you: The effect of social partner on cortisol and behavior during adolescence in a female rodent.” Ethology 130 e13414. doi: 10.1111/eth.13414.
- von Kortzfleisch, VT, and Richter, SH. . “Systematic heterogenization revisited: Increasing variation in animal experiments to improve reproducibility?” Journal of Neuroscience Methods 401 109992. doi: 10.1016/j.jneumeth.2023.109992.
- Kaiser, M, Gadau, J, Kaiser, S, Müller, C, and Richter, SH. . “Individualized social niches in animals: Theoretical clarifications and processes of niche change.” BioScience 74: 146–158. doi: 10.1093/biosci/biad122.
- Schiller D, Yu ANC, Alia-Klein N, ..., Richter SH, ... Lowe L. . “The human affectome.” Neuroscience and Biobehavioral Reviews 158 105450. doi: 10.1016/j.neubiorev.2023.105450.
- Rystrom, TL, Richter, SH, Sachser, N, and Kaiser, S. . “Social niche shapes social behavior and cortisol concentrations during adolescence in female guinea pigs.” Hormones and Behavior 162 (105539). doi: 10.1016/j.yhbeh.2024.105539.
- Kaiser, M.I., Killin, A., Abendroth, A.-K., Back, M.D., Baune, B.T., Bilstein, N., Breitmoser, Y., Caspers, B.A., Gadau, J., Gossmann, T.I., Kaiser, S., Krüger, O., Kurtz, J.Lengersdorf, D., Malsch, A. K. F., Müller, C. Rauthmann, J.F., Reinhold, K., Richter, S.H., Stummer, C., Trappes, R., Voelcker-Rehage, C., and Wittmann, M.J. . “Individualisation and individualised science across disciplinary perspectives.” European Journal for Philosophy of Science 14 (41) 41. doi: 10.1007/s13194-024-00602-8.
- Sroka, MGU, Ambree, O, Dohmen, C, Palme, R, Kaiser, S, and Richter, SH. . “Personality matters – The interplay between consistent individual differences and mouse welfare in female C57BL6/J mice.” Frontiers in Animal Science 5 (14238). doi: 10.3389/fanim.2024.1423814.
- Göbel, M., Kaiser, S., Richter, S.H., Wagner, P., and Wehrenberg, G. . “Specifically Unspecific – Simple Devices as Feasible and Effective Enrichment for Small Mammals.” Animal Behavior and Cognition 11 (3): 305–329. doi: 10.26451/abc.11.03.05.2024.
- Stieger, B, Wesseler, Y, Kaiser, S, Sachser, N, and Richter, SH. . “Behavioral lateralization of mice varying in serotonin transporter genotype.” Frontiers in Behavioral Neuroscience 16 1095567. doi: 10.3389/fnbeh.2022.1095567.
- Kaiser, S, Korte, A, Wistuba, J, Baldy, M, Wissmann, A, Dubičanac, A, Richter, SH, and Sachser, N. . “Effects of castration and sterilization on cortisol responsiveness – a case study in male guinea pigs.” Frontiers in Veterinary Science 9 (1093157). doi: 10.3389/fvets.2022.1093157.
- Segelcke, D, Linneman, J, Pradier, B, Kronenberg, D, Stange, R, Richter, SH, Görlich, D, Baldini, N, Di Pompo, G, Verri, WAJr., Avnet, S, and Pogatzki-Zahn, EM. . “Behavioral voluntary and social bioassays enabling identification of complex and sex-dependent pain-(-related) phenotypes in rats with bone cancer.” Cancers 15 (1565). doi: 10.3390/cancers15051565.
- Quante, SM, Siewert, V, Palme, R, Kaiser, S, Sachser, N, and SH, Richter. . “The power of a touch: Regular touchscreen training but not its termination affects hormones and behavior in mice.” Frontiers in Behavioral Neuroscience 17. doi: 10.3389/fnbeh.2023.1112780.
- Waterkamp, C, von Kortzfleisch, VT, Neu, C, Kitanovski, S, Richter, SH, and Hoffmann, D. . “Increasing information gain in animal research by improving statistical model accuracy.” contribution to the International Workshop on Mathematical Modeling and Scientific Computing 2022, München doi: 10.1088/1742-6596/2514/1/012021.
- Mutwill, AM, Schielzeth, H, Richter, SH, Kaiser, S, and Sachser, N. . “Conditional on the social environment? Roots of repeatability in hormone concentrations of male guinea pigs.” Hormones and Behavior 155 105423.
- Bohn, L, Bierbaum, L, Kästner, N, von Kortzfleisch, VT, Kaiser, S, Sachser, N, and Richter, SH. . “Structural enrichment for laboratory mice: exploring the effects of novelty and complexity.” Frontiers in Veterinary Science 10 1207332. doi: 10.3389/fvets.2023.1207332.
- Kaiser, S, and Richter, SH. . “Don’t worry, be happy? Ein verhaltensbiologischer Blick auf das Gefühlsleben der Tiere.” in Das unterschätzte Tier, edited by N Sachser, N Kästner and T Zimmermann. Hamburg: Rowohlt Taschenbuch Verlag.
- Stieger, B, Palme, R, Kaiser, S, Sachser, N, and Richter, SH. . “When left is right: The effects of paw preference training on behaviour in mice.” Behavioural Brain Research 430 (113929).
- Von Kortzfleisch, VT, Ambrée, O, Karp, NA, Meyer, N, Novak, J, Palme, R, Rosso, M, Touma, C, Würbel, H, Kaiser, S, Sachser, N, and Richter, SH. . “Do multiple experimenters improve the reproducibility of animal studies?” PLoS Biology 20(3) (e3001564). doi: 10.1371/journal.pbio.3001564.
- Rystrom, TL, Prawitt, RC, Richter, SH, Sachser, N, and Kaiser, S. . “Repeatability of endocrine traits and dominance rank in female guinea pigs.” Frontiers in Zoology 19 (1): 4.. doi: 10.1186/s12983-021-00449-2.
- Feige-Diller, J, Herrera-Rivero, M, Witten, A, Stoll, M, Kaiser, S, Richter, SH, and Sachser, N. . “The impact of varying food availability on gene expression in the liver: Testing the Match-Mismatch hypothesis.” Frontiers in Nutrition 9 (910762).
- Bračić, M, Bohn, L, Siewert, V, von Kortzfleisch, V, Schielzeth, H, Kaiser, S, Sachser, N, and Richter, SH. . “Once an Optimist, Always an Optimist? Studying Cognitive Judgment Bias in Mice.” Behavioral Ecology 33: 775–788.
- Herrera-Rivero, M, Bohn, L, Witten, A, Jüngling, K, Kaiser, S, Richter, SH, Stoll, M, and Sachser, N. . “Transcriptional profiles in the mouse amygdala after a cognitive judgment bias test largely depend on the genotype.” Frontiers in Molecular Neuroscience 15 1025389. doi: 10.3389/fnmol.2022.1025389.
- Jaric, I, Voelkl, B, Clerc, M, Schmid, MW, Novak, J, Rosso, M, Rufener, R, von Kortzfleisch, VT, Richter, SH, Buettner, M, Bleich, A, Amrein, I, Wolfer, DP, Touma, C, Sunagawa, S, and Würbel, H. . “The rearing environment persistently modulates mouse phenotypes from the molecular to the behavioural level.” PLoS Biology 20 (10) e3001837. doi: 10.1371/journal.pbio.3001837.
- Siewert, V, and Richter, SH. . “Always Looking on the Bright Side of Life?” forschung. Das Magazin der Deutschen Forschungsgemeinschaft 3 (4): 24–27.
- Jaric, I, Voelkl, B, Clerc, M, Schmid, MW, Novak, J, Rosso, M, Rufener, R, von Kortzfleisch, VT, Richter, SH, Buettner, M, Bleich, A, Amrein, I, Wolfer, DP, Touma, C, Sunagawa, S, and Würbel, H. . “The rearing environment persistently modulates mouse phenotypes from the molecular to the behavioural level.” PLoS Biology 20 (10). doi: 10.1371/journal.pbio.3001837.
- Feige-Diller, J, Palme, R, Kaiser, S, Sachser, N, and Richter, SH. . “The impact of varying food availability on health and welfare in mice: Testing the Match-Mismatch hypothesis.” Physiology and Behavior 228 (113193). doi: 10.1016/j.physbeh.2020.113193.
- Mutwill, AM, Schielzeth, H, Zimmermann, TD, Richter, SH, Kaiser, S, and Sachser, N. . “Individuality meets plasticity: Endocrine phenotypes across male dominance rank acquisition in guinea pigs living in a complex social environment.” Hormones and Behavior 131 104967. doi: 10.1016/j.yhbeh.2021.104967.
- Desjardins, E, Kurtz, J, Kranke, N, Lindeza, A, and Richter, SH. . “Beyond standardization: Improving external validity and reproducibility in experimental evolution.” BioScience biab008. doi: 10.1093/biosci/biab008.
- Stieger, B, Melotti, L, Quante, SM, Kaiser, S, Sachser, N, and Richter, SH. . “A step in the right direction: the effect of context, strain and sex on paw preference in mice.” Animal Behaviour 174: 21–30. doi: 10.1016/j.anbehav.2021.01.012.
- Krakenberg, V, Wewer, M, Palme, R, Kaiser, S, Sachser, N, and Richter, SH. . “Regular touchscreen training affects faecal corticosterone metabolites and anxiety-like behaviour in mice.” Behavioural Brain Research 401: 113080. doi: 10.1016/j.bbr.2020.113080.
- Bodden, C, Wewer, M, Kästner, N, Palme, R, Kaiser, S, Sachser, N, and Richter, SH. . “Not all mice are alike: Mixed-strain housing alters social behaviour.” Physiology and Behavior 228 (113220). doi: 10.1016/j.physbeh.2020.113220.
- Alexander, R, Aragon, OR, Bookwala, J, Cherbuin, N, Gatt, JM, Kahrilas, IJ, Kästner, N, Lawrence, A, Lowe, L, Morrison, RG, Mueller, SC, Nusslock, R, Papadelis, C, Polnaszek, KL, Richter, SH, Silton, RL, and Styliadis, C. . “The neuroscience of positive emotions and affect: Implications for cultivating happiness and wellbeing.” Neuroscience and Biobehavioral Reviews 121: 220–249. doi: 10.1016/j.neubiorev.2020.12.002.
- Melotti, L, Siestrup, S, Peng, M, Vitali, V, Dowling, D, von Kortzfleisch, VT, Bracic, M, Sachser, N, Kaiser, S, and Richter, SH. . “Individuality, as well as genetic background, affects syntactical features of courtship songs in male mice.” Animal Behaviour 180: 179–196.
- SH, Richter. . “Reproduzierbarkeit von Verhaltensdaten (Kapitel 8.3).” in Methoden der Verhaltensbiologie, edited by M Naguib and ET Krause. Berlin: Springer Spektrum.
- SH, Richter. . “Automated home-cage testing as a tool to improve reproducibility of behavioral research?” Frontiers in Neuroscience 14: 383..
- Feige-Diller, J, Krakenberg, V, Bierbaum, L, Seifert, L, Palme, R, Kaiser, S, Sachser, N, and Richter, SH. . “The effects of different feeding routines on welfare in laboratory mice.” Frontiers in Veterinary Science 6: 479.
- Von Kortzfleisch, VT, Karp, NA, Palme, R, Kaiser, S, Sachser, N, and Richter, SH. . “Improving reproducibility in animal research by splitting the study population into several ‘mini-experiments’.” Scientific Reports 10 (16579). doi: 10.1038/s41598-020-73503-4.
- Krakenberg, V, Siestrup, S, Palme, R, Kaiser, S, Sachser, N, and Richter, SH. . “Effects of different social experiences on emotional state in mice.” Scientific Reports 10 (15255). doi: 10.1038/s41598-020-71994-9.
- Richter, SH, and von Kortzfleisch, V. . “It is time for an empirically informed paradigm shift in animal research.” Nature Reviews Neuroscience 21: 660. doi: 10.1038/s41583-020-0370-7.
- Mutwill, AM, Zimmermann, TD, Hennicke, A, Richter, SH, Kaiser, S, and Sachser, N. . “Adaptive reshaping of the hormonal phenotype after social niche transition in adulthood.” Proceedings of the Royal Society B: Biological Sciences 287 20200667. doi: 10.1098/rspb.2020.0667.
- Krakenberg, Viktoria, Siestrup, Sophie, Palme, Rupert, Kaiser, Sylvia, Sachser, Norbert, and Richter, Sophie Helene. . “Effects of different social experiences on emotional state in mice.” Scientific Reports 10 (15255). doi: 10.1038/s41598-020-71994-9.
- Mutwill, AM, Zimmermann, TD, Reuland, C, Fuchs, S, Kunert, J, Richter, SH, Kaiser, S, and Sachser, N. . “High reproductive success despite queuing – Socio-sexual development of males in a complex social environment.” Frontiers in Psychology 10 2810. doi: 10.3389/fpsyg.2019.02810.
- Krakenberg, V, von Kortzfleisch, V, Kaiser, S, Sachser, N, and Richter, SH. . “Differential effects of serotonin transporter genotype on anxiety-like behavior and cognitive judgment bias in mice.” Frontiers in Behavioral Neuroscience 13: 263.
- Haaker, J, Maren, S, Andreatta, M, Merz, CJ, Richter, J, Richter, SH, Drexler, Shira S, Lange, MD, Jüngling, K, Nees, F, Seidenbecher, T, Fullana, MA, Wotjak, C, and Lonsdorf, TB. . “Making translation work: Harmonizing cross-species methodology in the behavioural neuroscience of Pavlovian fear conditioning.” Neuroscience & Biobehavioral Reviews 107: 329–345.
- Bodden, C, von Kortzfleisch, V, Karwinkel, F, Kaiser, S, Sachser, N, and Richter, SH. . “Heterogenising study samples across testing time improves reproducibility of behavioural data.” Scientific Reports 9: 8247..
- Von Kortzfleisch, V, Kästner, N, Prange, L, Kaiser, S, Sachser, N, and Richter, SH. . “Have I been here before? Effects of repeated testing and age on the results of behavioural tests.” Behavioural Brain Research 367: 143–148.
- Melotti, L, Kästner, N, Eick, AK, Schnelle, AL, Palme, R, Sachser, N, Kaiser, S, and Richter, SH. . “Can live with ‘em, can live without ‘em: pair housed male C57BL/6J mice show low aggression and increasing sociopositive interactions with age, but can adapt to single housing if separated.” Applied Animal Behaviour Science 214: 79–88.
- Kästner, N, Richter, SH, Urbanik, S, Kunert, J, Waider, J, Lesch, K-P, Kaiser, S, and Sachser, N. . “Brain serotonin deficiency affects female aggression.” Scientific Reports 9 (1366).
- Krakenberg, V, Woigk, I, Garcia, Rodriguez L, Kästner, N, Kaiser, S, Sachser, N, and Richter, SH. . “Technology or ecology? New tools to assess cognitive judgement bias in mice.” Behavioural Brain Research 362: 279–287. doi: 10.1016/j.bbr.2019.01.021.
- Richter, SH, and Hintze, S. . “From the individual to the population – and back again? Emphasising the role of the individual in animal welfare science.” Applied Animal Behaviour Science 212: 1–8.
- Bodden, C, Siestrup, S, Palme, R, Kaiser, S, Sachser, N, and Richter, SH. . “Evidence-based severity assessment: Impact of repeated versus single open-field testing on welfare in C57BL/6J mice.” Behavioural Brain Research 336: 261–268.
- Kästner, N, Richter, SH, Bodden, C, Palme, R, Kaiser, S, and Sachser, N. . “Varying social experiences in adulthood do not differentially affect anxiety-like behavior but stress hormone levels.” Frontiers in Behavioral Neuroscience 12: 72..
- SH, Richter. . “‘Never replicate a successful experiment’ - Gedanken über Unschärfe im Tierversuch.” in Unschärfe – der Umgang mit fehlender Eindeutigkeit, edited by S Freitag, M Geierhos, R Asmani and JI Haug.
- Kafkafi, N, Agassi, J, Chesler, EJ, Crabbe, JC, Crusio, WE, Eilam, D, Gerlai, R, Golani, I, Gomez-Marin, A, Heller, R, Iraqi, F, Jaljuli, I, Karp, NA, Morgan, H, Nicholson, G, Pfaff, DW, Richter, SH, Stark, PB, Stiedl, O, Stodden, V, Tarantino, LM, Tucci, V, Valdar, W, Williams, RW, Würbel, H, and Benjamini, Y. . “Reproducibility and replicability of rodent phenotyping in preclinical studies.” Neuroscience & Biobehavioral Reviews 87: 218–232.
- SH, Richter. . “Aus dem VBIO.” Biologie in unserer Zeit (BIUZ) 48: 23..
- Sachser, N, Richter, SH, and Kaiser, S. . “Artgerecht/tiergerecht.” in Handbuch Tierethik, edited by JS Ach and D Borchers. Stuttgart: J.B. Metzler Verlag.
- Richter, SH, Sachser, N, and Kaiser, S. . “Emotionen.” in Handbuch Tierethik, edited by JS Ach and D Borchers. Stuttgart: J.B. Metzler Verlag.
- SH, Richter. . “Systematic heterogenization for better reproducibility in animal experimentation.” Lab Animal 46: 343–349. doi: 10.1038/laban.1330.
- Meyer, N, Jenikejew, J, Richter, SH, Kaiser, S, and Sachser, N. . “Social experiences during adolescence affect anxiety-like behavior but not aggressiveness in male mice.” Behavioural Brain Research 326: 147–153. doi: 10.1016/j.bbr.2017.03.017.
- Kästner, N, Richter, SH, Gamer, M, Kaiser, S, and Sachser, N. . “What a difference a day makes - female behaviour is less predictable near ovulation.” Royal Society Open Science 4. doi: 10.1098/rsos.160998.
- Remmes, J, Bodden, C, Richter, SH, Lesting, J, Sachser, N, Pape, H-C, and Seidenbecher, T. . “Impact of life history on fear memory and extinction.” Frontiers in Behavioural Neuroscience 10: 185. doi: 10.3389/fnbeh.2016.00185.
- Graulich, DM, Kaiser, S, Sachser, N, and Richter, SH. . “Looking on the bright side of bias – validation of an affective bias test for laboratory mice.” Applied Animal Behaviour Science 181: 173–181.
- Richter, SH, Spinello, C, and Macrì, S. . “Improving external validity of experimental animal data.” in Animal Models for Human Cancer: Discovery and Development of Novel Therapeutics, edited by MI Martic-Kehl and PA Schubiger. Weinheim: Wiley-VCH Verlag.
- Mallien, AS, Palme, R, Richetto, J, Muzzillo, C, Richter, SH, Vogt, MA, Inta, D, Riva, MA, Vollmayr, B, and Gass, P. . “Daily exposure to a touchscreen-paradigm and associated food restriction evokes an increase in adrenocortical and neural activity in mice.” Hormones and Behavior 81: 97–105.
- Meyer, N, Richter, SH, Schreiber, RS, Kloke, V, Kaiser, S, Lesch, KP, and Sachser, N. . “The Unexpected Effects of Beneficial and Adverse Social Experiences during Adolescence on Anxiety and Aggression and Their Modulation by Genotype.” Frontiers in Behavioral Neuroscience 10 (97). doi: 10.3389/fnbeh.2016.00097.
- Richter, SH, Kästner, N, Loddenkemper, DH, Kaiser, S, Sachser, and N. . “A time to wean? Impact of weaning age on anxiety-like behaviour and stability of behavioural traits in full adulthood.” PloS one 11(12): e0167652.. doi: 10.1371/journal.pone.0167652.
- Richter, SH, Kästner, N, Kriwet, M, Kaiser, S, and Sachser, N. . “Play matters - the surprising relationship between juvenile playfulness and anxiety in later life.” Animal Behaviour 114: 261–271. doi: 10.1016/j.anbehav.2016.02.003.
- Sachser, N, and Richter, SH. . “Animals can tell us more.” Behavioral and Brain Sciences 38: 44–45.
- Macri, S, and Richter, SH. . “The Snark was a Boojum – reloaded.” Frontiers in Zoology 12: S20.
- Bodden, C, Richter, SH, Schreiber, RS, Kloke, V, Gerss, J, Palme, R, Lesch, K-P, Lewejohann, L, Kaiser, S, and Sachser, N. . “Benefits of adversity?! How life history affects the behavioral profile of mice varying in serotonin transporter genotype.” Frontiers in Behavioral Neuroscience 9: 47.
- Kästner, N., Richter, S.H., Lesch, K.P., Schreiber, R.S., Kaiser, S., and Sachser, N. . “Benefits of a "vulnerability gene"? A study in serotonin transporter knockout mice.” Behavioural Brain Research 283. doi: 10.1016/j.bbr.2015.01.031.
- Richter, SH, Vogel, AS, Ueltzhöffer, K, Muzzillo, C, Vogt, MA, Lankisch, K, Armbruster-Genç, DJ, Riva, MA, Fiebach, CJ, Gass, P, and Vollmayr, B. . “Touchscreen-paradigm for mice reveals cross-species evidence for an antagonistic relationship of cognitive flexibility and stability.” Frontiers in Behavioral Neuroscience 8: 154.. doi: 10.3389/fnbeh.2014.00154.
- Lima-Ojeda, JM, Vogt, MA, Richter, SH, Dormann, C, Schneider, M, Gass, P, and Inta, D. . “Lack of protracted behavioral abnormalities following intermittent or continuous chronic mild hypoxia in perinatal C57BL/6 mice.” Neuroscience Letters 577C: 77–82. doi: 10.1016/j.neulet.2014.06.022.
- Richter, SH, Wollmann, E, Schmidt, M, Zillmann, U, Hellweg, R, Sprengel, R, and Gass, P. . “The effects of neonatal cryoanaesthesia-induced hypothermia on adult emotional behaviour and stress markers in C57BL/6 mice.” Behavioural Brain Research 270C: 300–306. doi: 10.1016/j.bbr.2014.05.002.
- Richter, SH, Gass, P, and Fuss, J. . “Resting Is Rusting: A Critical View on Rodent Wheel-Running Behavior.” Neuroscientist 20(4): 313–325. doi: 10.1177/1073858413516798.
- Richter, SH, Sartorius, A, Gass, P, and Vollmayr, B. . “A matter of timing: harm reduction in learned helplessness.” Behavioral and Brain Functions 10: 41..
- Richter, SH, Zeuch, B, Lankisch, K, Gass, P, Durstewitz, D, and Vollmayr, B. . “Where have I been? Where should I go? Spatial working memory on a radial arm maze in a rat model of depression.” PloS one 8 (4): e62458.. doi: 10.1371/journal.pone.0062458.
- Hoyer, C, Richter, H, Brandwein, C, Riva, MA, and Gass, P. . “Preconceptional paternal exposure to a single traumatic event affects postnatal growth of female but not male offspring.” NeuroReport 24 (15): 856–60. doi: 10.1097/WNR.0000000000000007.
- Richter, SH, Zeuch, B, Riva, MA, Gass, P, and Vollmayr, B. . “Environmental enrichment ameliorates depressive-like symptoms in young rats bred for learned helplessness.” Behavioural Brain Research 252: 287–92. doi: 10.1016/j.bbr.2013.06.021.
- Würbel, H, Richter, SH, and Garner, JP. . “Reply to: "Reanalysis of Richter et al. (2010) on reproducibility".” Nature Methods 10 (5): 374.. doi: 10.1038/nmeth.2446.
- Fuss, J, Richter, SH, Steinle, J, Deubert, G, Hellweg, R, and Gass, P. . “Are you real? Visual simulation of social housing by mirror image stimulation in single housed mice.” Behavioural Brain Research 243: 191–8. doi: 10.1016/j.bbr.2013.01.015.
- Pignatelli, M, Vollmayr, B, Richter, SH, Middei, S, Matrisciano, F, Molinaro, G, Nasca, C, Battaglia, G, Ammassari-Teule, M, Feligioni, M, Nisticò, R, Nicoletti, F, and Gass, P. . “Enhanced mGlu5-receptor dependent long-term depression at the Schaffer collateral-CA1 synapse of congenitally learned helpless rats.” Neuropharmacology 66: 339–47. doi: 10.1016/j.neuropharm.2012.05.046.
- Richter, SH, Schick, A, Hoyer, C, Lankisch, K, Gass, P, and Vollmayr, B. . “A glass full of optimism: enrichment effects on cognitive bias in a rat model of depression.” Cognitive, Affective, and Behavioral Neuroscience 12 (3): 527–42. doi: 10.3758/s13415-012-0101-2.
- Gross, AN, Richter, SH, Engel, AK, and Würbel, H. . “Cage-induced stereotypies, perseveration and the effects of environmental enrichment in laboratory mice.” Behavioural Brain Research 234 (1): 61–8. doi: 10.1016/j.bbr.2012.06.007.
- Hoyer, C, Vogt, MA, Richter, SH, Zaun, G, Zahedi, Y, Maderwald, S, Ladd, ME, Winterhager, E, Grümmer, R, and Gass, P. . “Repetitive exposure to a 7 Tesla static magnetic field of mice in utero does not cause alterations in basal emotional and cognitive behavior in adulthood.” Reproductive Toxicology 34 (1): 86–92. doi: 10.1016/j.reprotox.2012.03.006.
- Chourbaji, S, Hoyer, C, Richter, SH, Brandwein, C, Pfeiffer, N, Vogt, MA, Vollmayr, B, and Gass, P. . “Differences in mouse maternal care behavior - is there a genetic impact of the glucocorticoid receptor?” PloS one 6 (4): e19218.. doi: 10.1371/journal.pone.0019218.
- Richter, SH, Garner, JP, Zipser, B, Lewejohann, L, Sachser, N, Touma, C, Schindler, B, Chourbaji, S, Brandwein, C, Gass, P, van Stipdonk, N, van der, Harst J, Spruijt, B, Võikar, V, Wolfer, DP, and Würbel, H. . “Effect of population heterogenization on the reproducibility of mouse behavior: a multi-laboratory study.” PloS one 6 (1): e16461.. doi: 10.1371/journal.pone.0016461.
- Gross, AN, Engel, AK, Richter, SH, Garner, JP, and Würbel, H. . “Cage-induced stereotypies in female ICR CD-1 mice do not correlate with recurrent perseveration.” Behavioural Brain Research 216 (2): 613–20. doi: 10.1016/j.bbr.2010.09.003.
- SH, Richter. . “Illusion der perfekten Kontrolle.” Bild der Wissenschaft plus 2011: 20–23.
- SH, Richter. . “Illusion der perfekten Kontrolle.” Vet Journal 12/11: 46–49.
- Engel, AK, Gross, AN, Richter, SH, and Würbel, H. . “Variation in stress reactivity affects cage-induced stereotypies in female CD-1 (ICR) mice.” Applied Animal Behaviour Science 133: 101–108.
- Richter, SH, Garner, JP, Auer, C, Kunert, J, and Würbel, H. . “Systematic variation improves reproducibility of animal experiments.” Nature Methods 7 (3): 167–8. doi: 10.1038/nmeth0310-167.
- SH, Richter. . “Standardization in animal experiments: Necessity or fallacy?” Dissertation thesis, Universität Münster.
- Richter, SH, Garner, JP, and Würbel, H. . “Environmental standardization: cure or cause of poor reproducibility in animal experiments?” Nature Methods 6 (4): 257–61. doi: 10.1038/nmeth.1312.
- Ambrée, O, Richter, H, Sachser, N, Lewejohann, L, Dere, E, de Souza, Silva MA, Herring, A, Keyvani, K, Paulus, W, and Schäbitz, WR. . “Levodopa Ameliorates Learning and Memory Deficits in a Murine Model of Alzheimer's Disease.” Neurobiology of Aging 30 (8): 1192–1204. doi: 10.1016/j.neurobiolaging.2007.11.010.
- Richter, H, Ambrée, O, Lewejohann, L, Herring, A, Keyvani, K, Paulus, W, Palme, R, Touma, C, Schäbitz, WR, and Sachser, N. . “Wheel-Running in a Transgenic Mouse Model of Alzheimer's Disease: Protection or Symptom?” Behavioural Brain Research 190 (1): 74–84. doi: 10.1016/j.bbr.2008.02.005.
- SH, Richter. . Was ist Glück?, edited by B Heller.