Dr. rer. nat. Harald Kullmann

• Evolutions- und Verhaltensökologie
• Evolution von Paarungssystemen (Buntbarsche, Skorpionsfliegen)
• Evolution von Karyotypen bei Killifischen

Academic Education

1994 – 1998

Promotion am Institut für Evolutionsbiologie und Ökologie, Universität Bonn

1988 – 1994

Studium der Biologie an der Universität Bonn

Positions

since 2008

Wissenschaftlicher Mitarbeiter am Zentrum für Didaktik der Biologie, Münster

1998 – 2008

Wissenschaftlicher Assistent am Institut für Evolutionsbiologie und Ökologie, Universität Bonn

Summer Term 2026

• V/Ü: Naturwissenschaften im Zusammenhang [134024]
  (in cooperation with Birte Müller and Anna Bröker)
  • [17^th Apr | Fri, 12.00 p. m. – 1.00 p. m. | weekly | ZH]
  • [21^st Apr | Tue, 2.00 p. m. – 4.00 p. m. | weekly | SP4 108]
  • [21^st Apr | Tue, 12.00 p. m. – 2.00 p. m. | weekly | SP4 108]
  • [21^st Apr | Tue, 12.00 p. m. – 2.00 p. m. | weekly | SP4 109]
  • [21^st Apr | Tue, 4.00 p. m. – 6.00 p. m. | weekly | SP4 108]
  • [22^nd Apr | Wed, 8.00 a. m. – 10.00 a. m. | weekly | SP4 108]
  • [22^nd Apr | Wed, 12.00 p. m. – 2.00 p. m. | weekly | SP4 107]
  • [22^nd Apr | Wed, 12.00 p. m. – 2.00 p. m. | weekly | SP4 108]
  • [22^nd Apr | Wed, 2.00 p. m. – 4.00 p. m. | weekly | SP4 107]
  • [22^nd Apr | Wed, 4.00 p. m. – 6.00 p. m. | weekly | SP4 108]
  • [22^nd Apr | Wed, 4.00 p. m. – 6.00 p. m. | weekly | SP4 107]
  • [22^nd Apr | Wed, 8.00 a. m. – 10.00 a. m. | weekly | SP4 106]
  • [23^rd Apr | Thu, 8.00 a. m. – 10.00 a. m. | weekly | SP4 108]
  • [23^rd Apr | Thu, 4.00 p. m. – 6.00 p. m. | weekly | SP4 109]
  • [23^rd Apr | Thu, 2.00 p. m. – 4.00 p. m. | weekly | SP4 109]
  • [24^th Apr | Fri, 8.00 a. m. – 10.00 a. m. | weekly | SP4 106]
  • [24^th Apr | Fri, 10.00 a. m. – 12.00 p. m. | weekly | SP4 109]
  • [24^th Apr | Fri, 8.00 a. m. – 10.00 a. m. | weekly | SP4 108]
  • [24^th Apr | Fri, 8.00 a. m. – 10.00 a. m. | weekly | SP4 109]
  • [24^th Apr | Fri, 10.00 a. m. – 12.00 p. m. | weekly | SP4 108]
  • [Wed, 13^th May 2026, 8.00 a. m. – 10.00 a. m. | Single Course | SP4 109]
  • [Sat, 16^th May 2026, 10.30 a. m. – 12.30 p. m. | Single Course | F1]
  • [Wed, 3^rd Jun 2026, 8.00 a. m. – 10.00 a. m. | Single Course | SP4 109]
  • [ – 20^th Jun 2026 | Sat, 10.30 a. m. – 12.30 p. m. | weekly | F1]
  • [ – 18^th Jul 2026 | Sat, 10.30 a. m. – 12.30 p. m. | weekly | F1]
• V/Ü: *) Freilandbiologie, zoologischer Teil [134014]
  (in cooperation with Robert Boczki)
  • [13^th Apr | Mon, 6.00 p. m. – 8.00 p. m. | weekly | MB K8]
  • [13^th Apr | Mon, 4.00 p. m. – 6.00 p. m. | weekly | MB K8]
  • [13^th Apr | Mon, 2.00 p. m. – 4.00 p. m. | weekly | MB K8]
  • [13^th Apr | Mon, 12.00 p. m. – 1.00 p. m. | weekly | SP 7]
  • [14^th Apr | Tue, 6.00 p. m. – 8.00 p. m. | weekly | MB K8]
  • [14^th Apr | Tue, 4.00 p. m. – 6.00 p. m. | weekly | MB K8]
  • [14^th Apr | Tue, 2.00 p. m. – 4.00 p. m. | weekly | MB K8]
• Hands-on course: *) Praxisphase [134077]
  (in cooperation with Julia Ghelman, Maria Sieverdingbeck, Miriam Pott, Sascia Zielonka, Klaus Tenberge and Anna Bröker)

Winter Term 2025/26

• Lecture: Grundlagen der Naturwissenschaften [132023]
• Lecture: Einführung in die Biologie [132007]
• Practice: *) Lerngruppe Chemie [132040]
  (in cooperation with Birte Müller and Anna Bröker)
• Practice: *) Lerngruppe Physik [132026]
  (in cooperation with Birte Müller and Anna Bröker)
• Practice: *) Lerngruppe Biologie [132025]
  (in cooperation with Birte Müller and Anna Bröker)
• Alle: MAP Grundlagenmodul Naturwissenschaften

Summer Term 2025

• V/Ü: Naturwissenschaften im Zusammenhang [130021]
  (in cooperation with Birte Müller and Anna Bröker)
• V/Ü: *) Freilandbiologie, zoologischer Teil [130007]
  (in cooperation with Robert Boczki)
• Practical/Seminar: *) Die heimischen Lebensräume [130027]
  (in cooperation with Kai Müller, Elisabeth Meyer, Maik Bartelheimer, Roman Asshoff, Hans-Uwe Schütz, Franz Goller and Joachim Kurtz)
• Alle: *) Thüringen-Exkursion mit Schwerpunkt "Außerschulische Lernorte" [130064]
  (in cooperation with Roman Asshoff)
• Hands-on course: *) Praxisphase [130017]
  (in cooperation with Anna Bröker, Maria Sieverdingbeck, Miriam Pott, Sascia Zielonka, Klaus Tenberge and Julia Ghelman)

• 2018
• 2016
• 2012
• 2011
• 2010
• 2009
• 2008
• 2007
• 2006
• 2005
• 1998

2018

• Jördens, J., Asshoff, R., Kullmann, H., & Hammann, M. (2018). Interrelating Concepts from Genetics and Evolution: Why are Cod Shrinking? The American Biology Teacher, 80 (2), 132–138. doi: 10.1525/abt.2018.80.2.132.

2016

• Jördens, J., Asshoff, R., Kullmann, H., & Hammann, M. (2016). Providing vertical coherence in explanations and promoting reasoning across levels of biological organization when teaching evolution. International Journal of Science Education, 38 (6), 960–992. doi: 10.1080/09500693.2016.1174790.

2012

• Kullmann, H., & Hammann, M. (2012). Evolution heute. Unterricht Biologie, 379, 2–10.
• Kullmann, H. (2012). Das Schrumpfen der Stoßzähne. Unterricht Biologie, 379 (379), 26–32.
• Kullmann, H., & Asshoff, R. (2012). Rasante Evolution: Parasit & Schmetterling. Unterricht Biologie, 379 (379), 33–38.

2011

• Langen, K., Schwarzer, J., Kullmann, H., Bakker, T., & Thünken, T. (2011). Microsatellite support for active inbreeding in a cichlid fish. PloS one, 6 (9), e24689.
• Thünken, T., Baldauf, S., Kullmann, H., Schuld, J., H., S., Bakker, & T. (2011). Size-related inbreeding preference and competitiveness in male Pelvicachromis taeniatus (Cichlidae). Behavioral Ecology, 22 (2), 358–362.
• Baldauf, S., B., T., Kullmann, H., Thünken, & T. (2011). Female nuptial coloration and its adaptive significance in a mutual mate choice system. Behavioral Ecology, 22 (3), 478–485.
• Jördens, J., Asshoff, R., Kullmann, H., Tyrrell, S., & Hammann, M. (2011). Situational interest in evolutionary topics, contexts and activities. in Yarden, A., & Carvalho, G. (eds.), Authenticity in Biology Education. Benefits and Challenges. A selection of papers presented at the VIIIth Conference of European Researchers in Didactics of Biology (ERIDOB). (pp. 225–236). Selbstverlag / Eigenverlag.
• Jördens, J., Asshoff, R., & Kullmann, H. (2011). Stammbäume lesen und verstehen. in Dreesmann, D., & Graf, D. u. (ed.), Evolutionsbiologie – Moderne Themen für den Unterricht (1. edt.) (pp. 377–403). Spektrum Akademischer Verlag.

2010

• Thünken, T., Meuthen, D., Bakker, T., & Kullmann, H. (2010). Parental investment in relation to offspring quality in the biparental cichlid fish Pelvicachromis taeniatus. Animal Behaviour, 07/2010.
• Thünken, T., Baldauf, S., Bersau, N., Bakker, T., Kullmann, H., & Frommen, J. (2010). Impact of olfactory non-host predator cues on aggregation behaviour and activity in Polymorphus minutus infected Gammarus pulex. Hydrobiologia, 654 (1), 137–145.
• Baldauf, S., Bakker, T., Herder, F., K., H., Thünken, & T. (2010). Male mate choice scales female ornament allometry in a cichlid fish. BMC Evolutionary Biology, 301 (10).

2009

• Baldauf, S., Kullmann, H., Thünken, T., Winter, S., & Bakker, T. (2009). Computer animation as a tool to study preferences in the cichlid Pelvicachromis taeniatus. Journal of Fish Biology, 75 (3), 738–746.
• Kullmann, H., & Sauer, K.P. (2009). Mating tactic dependent sperm transfer rates in Panorpa similis (Mecoptera; Panorpidae): a case of female control? Ecological Entomology, 34 (1), 153–157.
• Thünken, T., Waltschyk, N., Bakker, T., & Kullmann, H. (2009). Olfactory self-recognition in a cichlid fish. Animal Cognition, 12 (5), 717–724.
• Baldauf, S., Kullmann, H., Schroth, S., Thünken, T., & Bakker, T. (2009). You can't always get what you want: size assortative mating by mutual mate choice as a resolution of sexual conflict. BMC Evolutionary Biology, 129 (9).
• Baldauf, S., Kullmann, H., Thünken, T., Winter, S., & Bakker, T. (2009). Computer animation as a tool to study preferences in the cichlid Pelvicachromis taeniatus. Journal of Fish Biology, 75 (3), 738–746.
• Kullmann, H., & Sauer, K.P. (2009). Mating tactic dependent sperm transfer rates in Panorpa similes (Mecoptera; Panorpidae): A case of female control? Ecological Entomology, 34, 153–157.

2008

• Völker, M., Ráb, P., & Kullmann, H. (2008). Karyotype differentiation in Chromaphyosemion killifishes (Cyprinodontiformes, Nothobranchiidae): Patterns, mechanisms, and evolutionary implications. Botanical Journal of the Linnean Society, 94 (1), 143–153.
• Baldauf, S., Kullmann, H., & Bakker, T. (2008). Technical Restrictions of Computer Manipulated Visual Stimuli and Display Units for Studying Animal Behaviour. Ethology, 114 (8), 737–751.
• Kullmann, H., Thünken, T., Baldauf, S., Bakker, T., & Frommen, J. (2008). Fish odour triggers conspecific attraction behaviour in an aquatic invertebrate. Biology Letters, 4 (5), 458–460.
• Völker, M., Ráb, P., & Kullmann, H. (2008). Karyotype differentiation in Chromaphyosemion killifishes (Cypronodontiformes; Nothobranchiidae): Patterns, mechanisms and evolutionary implications. Botanical Journal of the Linnean Society, 94, 143–153.
• Kullmann, H., Thünken, T., Baldauf, S., Bakker, T.C.M., & Frommen, J.G. (2008). Fish odour triggers conspecific attraction behaviour in an aquatic invertebrate. Biology Letters, 4 (5), 458–460.

2007

• Volker, M., Sonnenberg, R., Ráb, P., & Kullmann, H. (2007). Karyotype differentiation in Chromaphyosemion killifishes (Cyprinodontiformes, Nothobranchiidae). III: extensive karyotypic variability associated with low mitochondrial haplotype differentiation in C. bivittatum. Cytogenetic and Genome Research, 116 (1-2), 116–126.
• Baldauf, A., Thünken, T., Frommen, J., Bakker, T., Heupel, O., & Kullmann, H. (2007). Infection with an acanthocephalan manipulates an amphipod's reaction to a fish predator's odours. International Journal for Parasitology, 37 (1), 61–65.
• Kullmann, H., & Klemme, B. (2007). Female mating preference for own males on species and population level in Chromaphyosemion killifishes (Cyprinodontiformes, Nothobranchiidae). Zoology, 110 (5), 377–386.
• Bakker, T., Lubjuhn, T., Kullmann, H., & Starck, J.M. (2007). Causal and historical evolutionary biology: papers in honour of Professor Klaus Peter Sauer. Zoology, 110 (5), 331–335.
• Völker, M., Ráb, P., & Kullmann, H. (2007). Banded karyotypes of Chromaphyosemion poliaki and C. volcanum (Cyprinodontoformes, Nothobranchiidae), with a discussion of the validity of C. poliaki. Ichthyological Exploration of Freshwaters, 18 (8), 1–8.
• Thünken, T., Bakker, T., Baldauf, S., & Kullmann, H. (2007). Active inbreeding in a cichlid fish and its adaptive significance. Current biology, 17 (3), 225–229.
• Thünken, T., Bakker, T., & Kullmann, H. (2007). Extraordinarily long sperm in the socially monogamous cichlid fish Pelvicachromis taeniatus. Naturwissenschaften, 94 (6), 489–491.
• Thünken, T., Bakker, T., Baldauf, S., & Kullmann, H. (2007). Direct Familiarity Does Not Alter Mating Preference for Sisters in Male Pelvicachromis taeniatus (Cichlidae). Ethology, 113 (11), 1107–1112.
• Sauer, K.P., & Kullmann, H. (2007). Die Entdeckung der Evolution. in Höxtermann, E., & Hilger, H. (eds.), Lebenswissen – Eine Einführung in die Geschichte der Biologie. (pp. 244–273). Natur+Text.
• Thünken, T., Baldauf, S., Bakker, T.C.M., & Kullmann, H. (2007). Active inbreeding in a cichlid fish and its adaptive significance. Current biology, 17, 225–229.
• Schmitt-Scheersoi, A., & Kullmann, H. (2007). Gendrift und Selektion spielerisch vermitteln. Praxis der Naturwissenschaften - Biologie in der Schule, 56 (7), 45–47.
• Kullmann, H., & Scheersoi, A. (2007). Sexuelle Selektion: Unterrichtsvorschlag. Praxis der Naturwissenschaften - Biologie in der Schule, 56 (5), 34–39.

2006

• Völker, M., & Kullmann, H. (2006). Sequential chromosome banding from single acetic acid fixed embryos of Chromaphyosemion killifishes (Cyprinodontiformes, Nothobranchiidae). Cybium: International journal of ichthyology, 30 (2), 171–176.
• Volker, M., Sonnenberg, R., Ráb, P., & Kullmann, H. (2006). Karyotype differentiation in Chromaphyosemion killifishes (Cyprinodontiformes, Nothobranchiidae). II: cytogenetic and mitochondrial DNA analyses demonstrate karyotype differentiation and its evolutionary direction in C. riggenbachi. Cytogenetic and Genome Research, 115 (1), 70–83.

2005

• Kullmann, H., & Sauer, K.P. (2005). Life histories and mating System aspects of two Caucasian scorpionfly species: Panorpa similis Esben-Petersen and Panorpa connexa Mac Lachlan. Zoologischer Anzeiger, 244 (1), 1–9.
• Volker, M., Sonnenberg, R., Ráb, P., & Kullmann, H. (2005). Karyotype differentiation in Chromaphyosemion killifishes (Cyprinodontiformes, Nothobranchiidae). I: Chromosome banding patterns of C. alpha, C. kouamense and C. lugens. Genetica, 125 (1), 33–41.

1998

• Sauer, K.P., Lubjuhn, T., Sindern, J., Kullmann, H., Kurtz, J., Epplen, C., & Epplen, J.T. (1998). Mating System and Sexual Selection in the Scorpionfly Panorpa vulgaris (Mecoptera: Panorpidae). Naturwissenschaften, 85 (5), 219–228.

2013

• Jördens, J.; Asshoff, R.; Kullmann, H.; Hammann, M. 2013: „Wirkung von Lernaufgaben zum evolutiven Wandel: Ebenenwechsel und Jo-Jo-Lernen“. Gesellschaft für Fachdidaktik e.V.: Lernaufgaben entwickeln, bearbeiten und überprüfen - Ergebnisse und Perspektiven der fachdidaktischen Forchung, Technische Universität Dortmund, Dortmund, Deutschland, Oct 8, 2013.

2011

• Jördens, Janina; Asshoff, Roman; Kullmann, Harald; Hammann, Marcus 2011: „Wirkung von Simulationen zu Variation und Selektion unter besonderer Berücksichtigung des Ebenenwechsels zwischen Phänotyp und Genotyp“. Internationale Tagung der FDdB im VBio: Didaktik der Biologie - Standortbestimmung und Perspektiven, Universität Bayreuth, Bayreuth, Deutschland, Sep 15, 2011.

2010

• Jördens, Janina; Asshoff, R.; Kullmann, H.; Hammann, M. 2010: “Interactions between student conceptions and teaching materials on evolution of life”. VIIIth Conference of European Researchers in Didactics of Biology, Universidade do Minho, Braga, Portugal, Jul 13, 2010.

2009

• Jördens, Janina; Asshoff, R.; Kullmann, H.; Hammann, M. 2009: „"Evolution of life" - Design und Evaluation von Unterrichtsmaterialien“. Internationale Tagung der FDdB im VBio: Heterogenität erfassen - individuell fördern im Biologieunterricht, Leibniz-Institut für die Pädagogik der Naturwissenschaften an der Universität Kiel, Kiel, Deutschland, Sep 24, 2009.