© N. Hölzel

Prof. Dr. Dr. h.c. Norbert Hölzel 

Biodiversity and Ecosystem Research Group
Institute of Landscape Ecology
Heisenbergstr. 2, D-48149 Münster
room 501
phone +49(0)251-83 33 994
fax +49(0)251-83 38 338
e-mail norbert.hoelzel [at] uni-muenster.de

office hours Tuesday 13-15 h and on appointment

  • Curriculum vitae

    since 10/2007 Full Professor for Ecosystem Research
    University of Münster, Institute of Landscape Ecology
    2005 - 2007 Visiting Professor for Ecosystem Research
    University of Münster, Institute of Landscape Ecology
    2004 Habilitation and venia legendi for Vegetation Ecology and Landscape Ecology
    Justus-Liebig-University Gießen, Faculty of Agriculture, Nutrition Science and Environmental Management
    Thesis Ecological significance of seed banks, germination and establishment for the restoration of flood-meadows
    1998 - 2004 Assistant Professor
    Justus-Liebig-University Gießen, Department of Landscape Ecology and Resources Management
    1996 - 1998 Research Associate
    Justus-Liebig-University Gießen, Department of Landscape Ecology and Resources Management
    1995 PhD in Geobotany
    Ludwig-Maximilians-University Munich, Faculty of Forest Sciences
    Thesis Erica pine forests in the northern limestone Alps
    1990 - 1996 free lance surveys
    numerous free lance surveys in vegetation ecology and nature conservation management for governmental agencies and private consulting enterprises
    1990 MS in Physical Geography
    Ludwig-Maximilians-University Munich, Department of Geography
    Thesis Vegetation development at land slides in the Bavarian Prealps
    1984 - 1990 Studies in Physical Geography, Soil Science, Landscape Ecology and Geobotany
    Ludwig-Maximilians-University Munich


  • Teaching

    • lectures: landscape ecology, landscapes of the world, geobotany, restoration ecology, soil ecology and site conditions
    • courses: multivariate statistical methods in vegetation and landscape ecology, scientific english
    • field courses: landscape ecology, restoration ecology, vegetation science, soil science, community ecology, physical geography/landscape ecology (for geography students)
    • field trips: Werdenfelser Land (2008), northwest Germany (Niederrhein, Recker Moor, Diepholzer Moorniederung, Baumberge)
    • students research projects: more
    • supervision of Diploma, MSc, BSc and state examination thesis as well as PhD thesis
  • Research

    • Grassland ecology (restoration and sustainable management of species-rich grasslands)
    • Floodplain ecology (nutrient gradients, response to flooding dynamics, dispersal processes, recruitment and vegetation patterns)
    • Ecology of rare and endangered species (habitat characteristics, population dynamics)
    • Seed ecology (phenology, soil seed banks and recruitment)
    • Restoration ecology (development and evaluation of techniques for the restoration of habitats and species)
    • Effects of Global Change (climate, land use) on landscapes, habitats and species
    • Development of adaptation and mitigation strategies for resource management, nature protection and biodiversity conservation
    • Land use change and landscape dynamics in Western Siberia and Kazakhstan
    • Main Research Topics


    Services, Memberships, Reviews, Awards


     Department of Geosciences, University of Münster

    • Faculty Council: full member since 10/2008
    • Chair person of the commission for research and young academics since 01/2009
    • Deputy Managing Director of the Institute of Landscape Ecology since 10/2008
    • Person in charge for the building AVZ Hüffergarten / Neubau Geo I 08/2008 - 01/2017



    Scientific Organisations


    Membership in scientific organisations

    • Society for Ecological Restoration International (SER), European Chapter
    • International Association of Vegetation Science (IAVS)
    • International Association for Landscape Ecology (IALE)
    • Ecological Society of Germany, Austria and Switzerland (GfÖ) (Working Groups Restoration Ecology and Population Ecology)
    • Floristisch-soziologische Arbeitsgemeinschaft (FlorSoz)
    • Reinhold-Tüxen-Gesellschaft (RTG)
    • Verband der Geographen an Deutschen Hochschulen (VDGH) (AK Biogeographie)
    • Deutscher Hochschulverband (DHV)
    • Arbeitsgemeinschaft Forstliche Standorts- und Vegetationskunde (AFSV)
    • Arbeitskreis Vegetationsdatenbanken
    • Botanische Vereinigung für Naturschutz in Hessen (BVNH)
    • Regensburgische Botanische Gesellschaft (RBG)



    • for international journals
      amongst others Acta Oecologica, Agriculture Ecosystems and Environment, Annals of Forest Science, Applied Vegetation Science, Basic and Applied Ecology, Biodiversity and Conservation, Biological Conservation, Ecography, Ecological Engineering, Ecosystems, Flora, Folia Geobotanica, Forest Ecology and Management, Global Change Biology, Journal for Nature Conservation, Journal of Applied Ecology, Journal of Ecology, Journal of Plant Nutrition and Soil Science, Journal of Vegetation Science, Nordic Journal of Botany, Phytocoenologia, Plant and Soil, Plant Biology, Plant Ecology, Polar Biology, Preslia, Remote Sensing of Environment, Restoration Ecology, Seed Science Research, Weed Research, Wetlands
    • for research foundations
      Deutsche Forschungsgemeinschaft DFG , VolkswagenStiftung, Czech Science Foundation



    • Honorary Doctorate by the University of Tyumen, Russia (04/2017)
    • Editors award ‘Paper of the Year’ by the chief editors of the journal Applied Vegetation Science for the article Restoration of a species-rich flood-meadow by topsoil removal and diaspore transfer with plant material by Hölzel & Otte (2003) [Editor's Award175 KB AVS]
  • Publications

    reprint request
    google scholar citations

    International publications (peer-reviewed)

    • Freitag M, Hölzel N, Neuenkamp L, van der Plas F, Manning P, Abrahão A, ... & Klaus V (2022) Increasing plant species richness by seeding has marginal effects on ecosystem functioning in agricultural grasslands. Authorea Preprints
    • Velbert F, Kleinebecker T, Klevesahl J, Schwartze P, Bucharova A, Hölzel N (submitted) Impact of different mowing regimes on the soil seed bank in wet grasslands – evidence from a 26 years field experiment
    • Wertebach T-M, Hölzel N, Kämpf I, Kiehl K, Knorr K-H, Kleinebecker T (submitted) Soil organic matter quality indicates a high potential for ongoing carbon sequestration in abandoned post-Soviet croplands
    • Apostolakis A, Schöning I, Klaus VH, Michalzik B, Bischoff WA, Boeddinghaus RS, …, Hölzel N, …, Schrumpf M (2022) Direct and plant community mediated effects of management intensity on annual nutrient leaching risk in temperate grasslands. Nutrient Cycling in Agroecosystems, 1-22 [doi: 10.1007/s10705-022-10209-1]

    • Heim RJ, Heim W, Bültmann H, Kamp J, Rieker D, Yurtaev A, Hölzel N (2022) Fire disturbance promotes biodiversity of plants, lichens and birds in the Siberian subarctic tundra. Global Change Biology, 28(3), 1048-1062 [doi: 10.1111/gcb.15963]

    • Heim RJ, Yurtaev A, Bucharova A, Heim W, Kutskir V, Knorr KH, ..., Hölzel N (2022) Fire in lichen-rich subarctic tundra changes carbon and nitrogen cycling between ecosystem compartments but has minor effects on stocks. Biogeosciences, 19(10), 2729-2740 [doi: 10.5194/bg-19-2729-2022]

    • Heim W, Thomas A, Berner I, Korschefsky T, Hölzel N, Kamp J (2022) Anthropogenic fire patterns affect niche breadth and niche overlap in sympatric songbird species. Science of The Total Environment, 833, 155160 [doi: 10.1016/j.scitotenv.2022.155160]

    • Lachaise T, Bergmann J, Hölzel N, Klaus VH, Kleinebecker T, Rillig MC, van Kleunen M (2022) Soil conditions drive below‐ground trait space in temperate agricultural grasslands. Journal of Ecology, 110(5), 1189-1200 [doi: 10.1111/1365-2745.13862]

    • Meyer UN, Tischer A, Freitag M, Klaus VH, Kleinebecker T, Oelmann Y, ..., Hölzel N, Hamer U (2022) Enzyme kinetics inform about mechanistic changes in tea litter decomposition across gradients in land-use intensity in Central German grasslands. Science of The Total Environment, 836, 155748 [doi: 10.1016/j.scitotenv.2022.155748]

    • Mudrák O, Hrabovský A, Dvorský M, Liancourt P, Albert AJ, Doležal J, …, Hölzel N, Májeková M (2022) Grassland restoration on former arable land: Fine‐scale grass accumulation and damaged soil conditions limit species establishment. Applied Vegetation Science, 25(2), e12665 [doi: 10.1111/avsc.12665]

    • Nguyen H, Trung TH, Phan DC, Anh Tran T, Thi Hai Ly N, Nasahara KN, ..., Hölzel N (2022) Transformation of rural landscapes in the Vietnamese Mekong Delta from 1990 to 2019: A spatio-temporal analysis. Geocarto International, 1-23 [doi: 10.1080/10106049.2022.2086623]

    • Rupprecht D, Jedrzejek B, Hölzel N (2022) Fallow deer foraging alone does not preserve the vegetation of traditionally sheep‐grazed calcareous grasslands. Journal of Applied Ecology [doi: 10.1111/1365-2664.14253]

    • Sikorski J, Baumgartner V, Birkhofer K, Boeddinghaus RS, Bunk B, Fischer M, Fösel BU, Friedrich MW, Göker M, Hölzel N, Huang S, Huber KJ,  Kandeler E, Klaus VH, Kleinebecker T, Marhan S, Von Mering C, Oelmann Y, Prati D, Regan KM, Richter-Heitmann T, Rodrigues JFM, Schmitt B, Schöning I, Schrumpf M, Schurig E, Solly EF, Wolters V, Overmann J (2022) The evolution of ecological diversity in Acidobacteria. Frontiers in Microbiology 78 [doi: 10.3389/fmicb.2022.715637]

    • Stroot L, Brinkert A, Hölzel N, Rüsing A, Bucharova A (2021) Establishment of wildflower strips in a wide range of environments: a lesson from a landscape‐scale project. Restoration Ecology, e13542 [doi: 10.1111/rec.13542]

    • Freitag M, Kamp J, Dara A, Kuemmerle T, Sidorova TV, Stirnemann IA, Velbert F, Hölzel N (2021) Post-Soviet shifts in fire and grazing regimes changed the functional plant community composition on the Eurasian steppe. Global Change Biology 27:388–401 [doi: 10.1111/gcb.15411]
    • Freitag M, Klaus VH, Bolliger R, Hamer U, Kleinebecker T, Prati D, Schäfer D, Hölzel N (2021) Restoration of plant diversity in permanent grassland by seeding: assessing the limiting factors along land-use gradients. Journal of Applied Ecology [doi: 10.1111/1365-2664.13883]
    • Heim RJ, Bucharova A, Brodt L, Kamp J, Rieker D, Soromotin AV, Yurtaev A, Hölzel N (2021) Post-fire vegetation succession in the Siberian subarctic tundra over 45 years. Science of The Total Environment 760: 143425 [doi: 10.1016/j.scitotenv.2020.143425]
    • Heim RJ, Heim W, Bültmann H, Kamp J, Rieker D, Yurtaev A, Hölzel N (2021) Fire disturbance promotes biodiversity at local and landscape scales in subarctic tundra. Global change biology [doi: 10.1111/gcb.15963]
    • Heim RJ, Heim W, Darman G, Heinken T, Smirenski SM, Hölzel N (2021) Litter removal through fire - a key process for wetland vegetation and ecosystem dynamics. Science of the Total Environment 755: 142659 [doi: 10.1016/j.scitotenv.2020.142659]
    • Heim W, Chan S, Hölzel N, Ktitorov P, Mischenko A, Kamp J (2021) East Asian buntings: ongoing illegal trade and encouraging conservation responses. Conservation Science and Practice 3: e405 [doi: 10.1111/csp2.405]
    • Heim W, Wobker J, Schmaljohann H (2021) Sex, age, molt strategy and migration distance explain the phenology of songbirds at a stopover along the East Asian flyway. Behavioural Ecology and Sociobiology 75 [doi:10.1007/s00265-020-02957-3]
    • Klein-Raufhake T, Höfner J, Hölzel N, Knorr KH, Lampei C, Mudrák O, Bucharova A (2021) Nitrogen limitation reduces the performance of target plant species in restored meadows. Restoration Ecology, e13608 [doi: 10.1111/rec.13608]
    • Le Provost G, Thiele J, Westphal C, Penone C, Allan E, Neyret M, van der Plas F, Ayasse M, Bardgett RD, Birkhofer K, Boch S, Bonkowski M, Buscot F, Feldhaar H, Gaulton R, Goldmann K, Gossner MM, Klaus VH, Kleinebecker T, Krauss J, Renner S, Scherreiks P, Sikorski J, Baulechner D, Blüthgen N, Bolliger R, Börschig C, Busch V, Chisté M, Fiore-Donno AM, Fischer M, Arndt H, Hölzel N, John K, Jung K, Lange M, Marzini C, Overmann J, Paŝalić E, Perović DJ, Prati D, Schäfer D, Schöning I, Schrumpf M, Sonnemann I, Steffan-Dewenter I, Tschapka M, Türke M, Vogt J, Wehner K, Weiner C, Weisser W, Wells K, Werner M, Wolters V, Wubet T, Wurst S, Zaitsev AS, Manning P (2021) Contrasting responses of above- and belowground diversity to multiple components of land-use intensity. Nature communications, 12(1), 1-13 [doi: 10.1038/s41467-021-23931-1]
    • Neff F, Brändle M, Ambarlı D, Ammer C, Bauhus J, Boch S, Hölzel N, Klaus VH, Kleinebecker T, Prati D, Schall P, Schäfer D, Schulze E-D, Seibold S, Simons NK, Weisser WW, Pellissier L, Gossner MM (2021) Changes in plant-herbivore network structure and robustness along land-use intensity gradients in grasslands and forests. Science Advances, 7(20), eabf3985 [doi: 10.1126/sciadv.abf3985]
    • Neyret M, Fischer M, Allan E, Hölzel N, Klaus VH, Kleinebecker T, Krauss J, Le Provost G, Peter S, Schenk N, Simons NK, van der Plas F, Binkenstein J, Börschig C, Jung K, Prati D, Schäfer D, Schäfer M, Schöning I, Schrumpf M, Tschapka M, Westphal C, Manning P (2021). Assessing the impact of grassland management on landscape multifunctionality. Ecosystem Services, 52, 101366 [doi: 10.1101/2020.07.17.208199]
    • Oelmann Y, Lange M, Leimer S, Roscher, Aburto F, Alt F, Bange N, BernerD, Boch S, Boeddinghaus RS, Buscot F, Dassen S, de Deyn G, Eisenhauer N, Gleixner G, Goldmann K, Hölzel N, Jochum M, Kandeler E, Klaus VH, Kleinebecker T, LeProvost G, Manning P, Marhan S, Prati D, Schäfer D, Schöning I, Schrumpf M, Schurig E, Wagg C, Wubet T, Wilcke W (2021). Above- and belowground biodiversity jointly tighten the P cycle in agricultural grasslands. Nature Communications, 12(1) [doi: 10.1038/s41467-021-24714-4]
    • Rupprecht D, Hölzel N, Bucharova A (2021): Is there local adaptation in plant species to soil reaction? A lesson from a multispecies experiment. Restoration Ecology [doi: 10.1111/rec.13393]
    • Sabatini FM, Lenoir J, Hattab T, Arnst EA, Chytrý M, Dengler J, De Ruffray P, [...], Hölzel N, [...], Kattge J (2021) sPlotOpen – An environmentally balanced, open-access, global dataset of vegetation plots. Global Ecology and Biogeography 30(9): 1740-1764 [doi:10.1111/geb.13346]
    • Těšitel J, Tahadlová M, Lepš J, Hölzel N (2021) Linking insect herbivory with plant traits: Phylogenetically structured trait syndromes matter. Journal of Vegetation Science, 32(4), e13061 [doi: 10.1111/jvs.13061]
    • Baumann M, Kamp J, Pötzschner F, Bleyhl B, Dara A, Hankerson B, Prishchepov AV, Schierhorn F, Müller D, Hölzel N, Krämer R, Urazaliyev R, Kuemmerle T (2020) Declining human pressure and opportunities for rewilding in the steppes of Eurasia. Diversity and Distributions, 26(9), 1058-1070 [doi: 10.1111/ddi.13110] Open Access
    • Dara A, Baumann M, Hölzel N, Hostert P, Kamp J, Müller D, Ullrich B, Kuemmerle T (2020) Post-Soviet land-use change affected fire regimes on the Eurasian steppes. Ecosystems 23, 943–956 [doi:10.1007/s10021-019-00447-w]
    • Dara A, Baumann M, Freitag M, Hölzel N, Hostert P, Kamp J, Müller D, Prishchepov AV, Kuemmerle T (2020) Annual Landsat time series reveal post-Soviet changes in grazing pressure. Remote Sensing of Enviroment 239: 111667 [doi:10.1016/j.rse.2020.111667]
    • Felipe-​Lucia M, Soliveres S, Penone C, Fischer M, Ammer C, Boch S, Boeddinghaus RS, Bonkowski M, Buscot F, Fiore-​Donno AM, Frank K, Goldmann K, Gossner MM, Hölzel N, Jochum M, Kandeler E, Klaus VH, Kleinebecker T, Leimer S, Manning P, Oelmann Y, Saiz H, Schall P, Schloter M, Schöning I, Schrumpf M, Solly EF, Stempfhuber B, Weiser WW, Wilcke W, Wubet T, Allan E (2020) Land-​use intensity alters biodiversity - ecosystem function - service networks. Proceedings of the National Academy of Sciences of the United States of America: 117: 28140-28149 [doi: 10.1073/pnas.2016210117]
    • Heim W, Heim RJ, Beermann I, Burkovskiy OA, Gerasimov Y, Ktitorov P, Ozaki K, Panov I, Sander MM, Sjöberg S, Smirenski SM, Thomas A, Tottrup A, Tiunov IM, Willemoes M, Hölzel N, Thorup K, Kamp J (2020) Using geolocator tracking data and ringing archives to validate citizen-science based seasonal predictions of songbird distribution in a data-poor region. Global Ecology and Conservation 24: e01215 [doi: 10.1016/j.gecco.2020.e01215]
    • Jochum M, Fischer M, Isbell F, Roscher C, van der Plas F, Boch S, Boenisch G, Buchmann N, Catford J, Cavender-​Bares J, Ebeling A, Eisenhauer N, Gleixner G, Hölzel N, Kattge J, Klaus VH, Kleinebecker T, Lange M, Le Provost G, Meyer ST, Molina-​Venegas R, Mommer L, Oelmann Y, Penone C, Prati D, Reich PB, Rindisbacher A, Schäfer D, Scheu S, Schmid B, Tilman D, Tscharntke T, Vogel A, Wagg C, Weigelt A, Weisser WW, Wilcke W, Manning P (2020) The results of biodiversity-​ecosystem functioning experiments are realistic. Nature Ecology & Evolution [doi: 10.1038/s41559-​020-1280-9]
    • Kamp J, Freitag M, Hölzel N (2020) Towards more meaningful scenarios of biodiversity responses to land-use change in Central Asia. Regional Environmental Change 20, 79 [doi:10.1007/s10113-020-01666-x]
    • Koshkina A, Grigoryeva I, Tokarsky V, Urazaliyev R, Kuemmerle T, Hölzel N, Kamp J (2020) Marmots from space: assessing population size and habitat use of a burrowing mammal using publicly available satellite images. Remote Sensing in Ecology and Conservation  6: 153-167 [doi:10.1002/rse2.138] Open Access
    • Pichon N, Cappelli S, Soliveres S, Hölzel N, Klaus VH, Kleinebecker T, Allan E (2020) Decomposition disentangled: a test of the multiple mechanisms by which nitrogen enrichment alters litter decomposition. Functional Ecology 34 (7), 1485-1496 [doi:10.1111/1365-2435.13560]
    • Boeddinghaus R, Marhan S, Berner D, Boch S, Fischer M, Hölzel N, Kattge J, Klaus VH, Kleinebecker T, Oelmann Y, Prati D, Schäfer D, Schöning I, Schrumpf M, Sorkau E, Kandeler E, Manning P (2019) Plant functional trait shifts explain concurrent changes in the structure and function of grassland soil microbial communities. Journal of Ecology 107: 2197–2210 [doi:10.1111/1365-2745.13182]
    • Bruelheide H,Dengler J,Jiménez-Alfaro B,Purschke O,Hennekens S, Chytrý M, Pillar V, Jansen F, Kattge J, Sandel B, Aubin I, Biurrun I, Field R, Haider S, Jandt U, Lenoir J, Peet R, Peyre G, Sabatini F, Schmidt M, Schrodt F, Winter M, Aćić S, Agrillo E, Alvarez M, Ambarli D, Angelini P, Apostolova I, Arfin Khan M, Arnst E, Attorre F, Baraloto C, Beckmann M, Berg C, Bergeron Y, Bergmeier E, Bjorkman A, Bondareva V, Borchardt P, Botta-Dukat Z, Boyle B, Breen A, Brisse H, Byun C, Cabido M, Casella L, Cayuela L, Černý T, Chepinoga V, Csiky J, Curran M, Cušterevska R, Dajić Stevanović Z, De Bie E, de Ruffray P, De Sanctis M, Dimopoulos P, Dressler S, Ejrnaes R, El Sheikh M, Enquist B, Ewald J, Fagundez J, Finckh M, Font X, Forey E, Fotiadis G, de Gasper A, Golub, Gutierrez A, Hatim MZ, He T, Higuchi P, Michalcova D, Hölzel N, Homeier J, Indreica A, Isık Gürsoy D, Jansen S, Janssen J, Jedrzejek B, Jiroušek M, Jürgens N, Kącki Z, Kavgacı A, Kearsley E, Kessler M, Knollova I, Kolomiychuk V, Korolyuk A, Kozhevnikova M, Kozub Ł, Krstonošić D, Kühl H, Kühn I, Kuzemko A, Küzmič F, Landucci F, Lee M, Levesley A, Li CF, Liu H, Lopez Gonzalez G, Lysenko T, Macanović A, Mahdavi P, Manning P, Marceno C, Martynenko V, Mencuccini M, Minden V, Moeslund J, Moretti M, Müller JV, Munzinger J, Niinemets U, Nobis M, Noroozi J, Nowak A, Onyshchenko V, Overbeck G, Ozinga W, Pauchard A, Pedashenko H, Penuelas J, Pérez-Haase A, Peterka T, Petrik P, Phillips O, Prokhorov V, Rašomavičius V, Revermann R, Rodwell J, Ruprecht E, Rūsina S, Samimi C, Schaminée J, Schmiedel U, Šibik J, Šilc U, Škvorc Z, Smyth A, Sop T, Sopotlieva D, Sparrow B, Svenning JC, Swacha G, Tang Z, Tsiripidis I, Turtureanu PD, Ugurlu E, Uogintas D, Valachovič M, Vanselow KA, Vashenyak Y, Vassilev K, Vélez-Martin E, Venanzoni R, Vibrans AC, Violle C, Virtanen R, von Wehrden H, Wagner V, Walker DA, Wana D, Weiher E, Wesche K, Whitfeld T, Willner W, Wiser S, Wohlgemuth T, Yamalov S, Zizka G, Zverev A (2019) sPlot – a new tool for global vegetation analyses. Journal of Vegetation Science 30: 161–186 [doi:10.1111/jvs.12710]
    • Bucharova A, Bossdorf O, Hölzel N, Kollmann J, Prasse R, Durka W (2019) Mix and match: regional admixture provenancing strikes a balance among different seed-sourcing strategies for ecological restoration. Conservation Genetics 20: 7–17 [doi:10.1007/s10592-018-1067-6]
    • Busch V, Klaus VH, Schäfer D, Prati D, Boch S, Müller J, Chisté M, Mody K, Blüthgen N, Fischer M, Hölzel N, Kleinebecker T (2019) Will I stay or will I go? Plant species specific response and tolerance to high land-use intensity in temperate grassland ecosystems. Journal of Vegetation Science 30: 674–686 [doi:10.1111/jvs.12749]
    • Dittberner H, Becker C, Jiao WB, Schneeberger K, Hölzel N, Tellier A, de Meaux J (2019) Strengths and potential pitfalls of hay-transfer for ecological restoration revealed by RAD-seq analysis in two floodplain Arabis species. Molecular Ecology 28(17): 3887–3091 [doi:10.1111/mec.15194]
    • Heim RJ, Hölzel N, Heinken T, Kamp J, Thomas A, Darman GF, Smirenski SM, Heim W (2019) Post-burn and long-term fire effects on plants and birds in floodplain wetlands of the Russian Far East. Biodiversity and Conservation 28: 1611–1628 [doi:10.1007/s10531-019-01746-3]


    © Springer
    • Kollmann J, Kirmer A, Tischew S, Hölzel N, Kiehl K, (eds.)(2019) Renaturierungsökologie, pp 1–491 [doi:10.1007/978-3-662-54913-1]
    • Midolo G, De Frenne P, Hölzel N, Wellstein C (2019) Global patterns of intraspecific leaf trait responses to elevation. Global Change Biology 25: 2485–2498 [doi:10.1111/gcb.14646]
    • Palpurina S, Chytrı M, Hölzel N, Tichý L, Wagner V, Horsák M, Axmanová I, Hajek M, Hájková P, Freitag M, Lososová Z, Mathar W, Tzonev R, Danihelka J, Dřevojan P (2019) The type of nutrient limitation affects the plant species richness–productivity relationship: evidence from dry grasslands across Eurasia. Journal of Ecology 107: 1038–1050 [doi:10.1111/1365-2745.13084]
    • Schäfer D, Klaus VH, Kleinebecker T, Boeddinghaus R, Hinderling J, Kandeler E, Marhan S, Nowak S, Sonnemann I, Wurst S, Fischer M, Hölzel N, Hamer U, Prati D (2019) Recovery of ecosystem functions after experimental disturbance in 73 grasslands differing in land-use intensity, plant species richness and community composition. Journal of Ecology 107(6): 2635–2649 [doi:10.1111/1365-2745.13211]
    • Schwabe A, Tischew S, Bergmeier E, Grave E, Härdtle W, Heinken T, Hölzel N, Peppler-Lisbach C, Remy D, Dierschke H (2019) Plant Community of the Year 2020: Mat grassland (Nardus stricta grassland) Tuexenia 39: 289–308 [doi:10.14471/2019.39.017]
    • van der Plas F, Allan E, Fischer M, Alt F, Arndt H, Binkenstein J, Blaser S, Blüthgen N, Böhm S, Hölzel N, Klaus VH, Kleinbecker T, Morris EK, Oelmann Y, Prati D, Renner S, Rillig MC, Schaefer HM, Schloter M, Schmitt B, Schöning I, Schrumpf M, Solly E, Sorkau E, Steckel J, Steffan-Dewenter I, Stempfhuber B, Tschapka M, Weiner CN, Weisser WW, Werner M, Westphal C, Wilcke W, Manning P (2019) Towards the development of general rules describing landscape heterogeneity-multifunctionality relationships. Journal of Applied Ecology 56: 168–179 [doi:10.1111/1365-2664.13260]
    • Vogt J, Klaus VH, Both S, Fürstenau C, Gockel S, Gossner MM, Heinze J, Hemp A, Hölzel N, Jung K, Kleinebecker T, Lauterbach R, Lorenzen K, Ostrowski A, Otto N, Prati D, Renner S, Schumacher U, Seibold S, Simons N, Steitz I, Teuscher M, Thiele J, Weithmann S, Wells K, Wiesner K, Ayasse M, Blüthgen N, Fischer M, Weisser WW (2019) Eleven years’ data of grassland management in Germany. Biodiversity Data Journal 7: e36387 [doi:10.3897/BDJ.7.e36387]


    • Bruelheide H, Dengler J, Purschke O, Lenoir J, Jiménez-Alfaro B, Hennekens SM, Botta-Dukát Z, Chytrý M, Field R, Jansen F, Kattge J, Pillar VD, Schrodt F, Mahecha MD, Peet RK, Sandel B, van Bodegom P, Altman J, Alvarez Davila E, Arfin Khan MAS, Attorre F, Aubin I, Baraloto C, Barroso JG, Bauters M, Bergmeier E, Biurrun I, Bjorkman AD, Blonder B, Čarni A, Cayuela L, Černý T, Cornelissen JHC, Craven D, Dainese M, Derroire G, De Sanctis M, Díaz S, Doležal J, Farfan-Rios W, Feldpausch TR, Fenton NJ, Garnier E, Guerin GR, Gutiérrez AG, Haider S, Hattab T, Henry G, Hérault B, Higuchi P, Hölzel N, Homeier J, Jentsch A, Jürgens N, Kącki Z, Karger DN, Kessler M, Kleyer M, Knollová I, Korolyuk AY, Kühn I, Laughlin DC, Lens F, Loos J, Louault F, Lyubenova MI, Malhi Y, Marcenò C, Mencuccini M, Müller JV, Munzinger J, Myers-Smith IH, Neill DA, Niinemets Ü, Orwin KH, Ozinga WA, Penuelas J, Pérez-Haase A, Petřík P, Phillips OL, Pärtel M, Reich PB, Römermann C, Rodrigues AV, Sabatini FM, Sardans J, Schmidt M, Seidler G, Silva Espejo JE, Silveira M, Smyth A, Sporbert M, Svenning J, Tang Z, Thomas R, Tsiripidis I, Vassilev K, Violle C, Virtanen R, Weiher E, Welk E, Wesche K, Winter M, Wirth C, Jandt U (2018) Global trait–environment relationships of plant communities. Nature Ecology and Evolution 2: 1907–1918 [doi:10.1038/s41559-018-0699-8]
    • Busch V, Klaus VH, Penone C, Schäfer D, Boch S, Prati D, Müller J, Socher S, Niinemets Ü, Peñuelas J, Hölzel N, Fischer M, Kleinebecker T (2018) Nutrient stoichiometry and land use rather than species richness determine plant functional diversity. Ecology and Evolution 8 (1) 601–616 [doi:10.1002/ece3.3609] Open Access
    • Dara A, Baumann M, Kuemmerle T, Pflugmacher D, Rabe A, Griffiths P, Hölzel N, Kamp J, Freitag M, Hostert P (2018) Mapping the timing of cropland abandonment and recultivation in northern Kazakhstan using annual Landsat time series. Remote Sensing of Environment 213: 49–60 [doi:10.1016/j.rse.2018.05.005]
    • Djukic I, Kepfer-Rojas S, Schmidt IK, Larsen KS, Beier C, Berg B, Verheyend K, TeaComposition (among others: Busch V, Hölzel N, Klaus VH, Kleinebecker T) (2018) Early stage litter decomposition across biomes. Science of the Total Environment 628-629: 1369–1394 [doi:10.1016/j.scitotenv.2018.01.012]
    • Elias D, Hölzel N, Tischew S (2018) Goat paddock grazing improves the conservation status of shrub-encroached dry grasslands. Tuexenia 38: 215–233 [doi:10.14471/2018.38.017]
    • Elias D, Hölzel N, Tischew S (2018) Positive effects of goat pasturing on the threatened spring geophyte Gagea bohemica in formerly abandoned dry grasslands. Flora 249: 53–59  [doi:10.1016/j.flora.2018.09.008 ]
    • Heer N, Klimmek F, Zwahlen C, Fischer M, Hölzel N, Klaus VH, Kleinebecker T, Prati D, Boch S (2018) Hemiparasite-density effects on grassland plant diversity, composition and biomass. Perspectives in Plant Ecology, Evolution and Systematics 32: 22–29 [doi:10.1016/j.ppees.2018.01.004]
    • Ivanova L, Yudina P, Ronzhina D, Ivanov L, Hölzel N (2018) Quantitative mesophyll parameters rather than whole-leaf traits predict response of C3 steppe plants to aridity. New Phytologist 217 (2): 558–570 [doi:10.1111/nph.14840]
    • Kamp J, Reinhard A, Frenzel M, Kämpfer S, Trappe J, Hölzel N (2018) Farmland bird responses to land abandonment in Western Siberia. Agriculture, Ecosystems and Environment 268: 61–69 [doi:10.1016/j.agee.2018.09.009]
    • Klaus VH, Hoever CJ, Fischer M, Hamer U, Kleinebecker T, Mertens D, Schäfer D, Prati D, Hölzel N (2018) Contribution of the soil seed bank to the restoration of temperate grasslands by mechanical sward disturbance. Restoration Ecology 26 (S2): 114–122 [doi:10.1111/rec.12626]
    • Klaus VH, Kleinebecker T, Busch V, Fischer M, Hölzel N, Nowak S, Prati D, Schäfer D, Schöning I, Schrumpf M, Hamer U (2018) Land use intensity, rather than plant species richness, affects the leaching risk of multiple nutrients from permanent grasslands . Global Change Biology 24 (7): 2828–2840 [doi:10.1111/gcb.14123]
    • Klaus VH, Schäfer D, Prati D, Busch V, Hamer U, Hoever CJ, Kleinebecker T, Mertens D, Fischer M, Hölzel N (2018) Effects of mowing, grazing and fertilization on soil seed banks in temperate grasslands in Central Europe. Agriculture, Ecosystems and Environment 256: 211–217 [doi:10.​1016/​j.​agee.​2017.​11.​008]
    • Kleinebecker T, Busch V, Hölzel N, Hamer U, Schäfer D, Prati D, Fischer M, Hemp A, Lauterbach R, Klaus VH (2018) And the winner is … ! A test of simple predictors of plant species richness in agricultural grasslands. Ecological Indicators 87: 296–301 [doi:10.1016/j.ecolind.2017.12.031]
    • Leimer S, Bischoff S, Boch S, Busch V, Escher P, Fischer M, Hänsel F, Hölzel N, Kerber K, VH Klaus, Kleinebecker T, Michalzik B, Nauss T, Schäfer D, Schöning I, Schwarz MT, Siemens J, Thieme L, Wöllauer S, Wilcke W (2018) Does plant diversity affect the water balance of established grassland systems? Ecohydrology 11 (4): e1945 [doi:10.1002/eco.1945]
    • Nguyen H, Hölzel N, Völker A, Kamp J(2018) Patterns and Determinants of Post-Soviet Cropland Abandonment in the Western Siberian Grain Belt. Remote Sensing 10 (12): 1973 [doi:10.3390/rs1012973] Open Access
    • Tischew S, Dierschke H, Schwabe A, Garve E, Heinken T, Hölzel N, Bergmeier E, Remy D, Härdtle W (2018) Plant Community of the Year 2019: Oatgras Meadow. Tuexenia 38: 287–295 [doi:10.14471/2018.38.011]


    • Bucharova A, Durka W, Hölzel N, Kollmann J, Michalski S, Bossdorf O (2017) Are local plants the best for ecosystem restoration? It depends on how you analyze the data. Ecology and Evolution 7 (24): 10683–10689 [doi:10.1002/ece3.3585]
    • Bucharova A, Michalski S, Hermann J-M, Heveling K, Durka W, Hölzel N, Kollmann J, Bossdorf O (2017) Genetic differentiation and regional adaptation among seed origins used for grassland restoration: lessons from a multispecies transplant experiment. Journal of Applied Ecology 54 (1): 127–136 [doi: 10.1111/1365-2664.12645]
    • Durka W, Michalski SG, Berendzen KW, Bossdorf O, Bucharova A, Hermann JM, Hölzel N, Kollmann J (2017) Genetic differentiation within multiple common grassland plants supports seed transfer zones for ecological restoration. Journal of Applied Ecology 54 (1): 116–126 [doi:10.1111/1365-2664.12636]
    • Gilhaus K, Freitag M, Kunze S, Hölzel N (2017) High fodder value and feeding likelihood favor endozoochorous plant dispersal. Journal of Vegetation Science 28 (2): 357–367 [doi:10.1111/jvs.12481]
    • Gilhaus K, Boch S, Fischer M, Hölzel, N, Kleinebecker T, Prati D, Rupprecht D, Schmitt B, Klaus VH (2017) Grassland management in Germany: effects on plant diversity and vegetation composition. Tuexenia 37: 379–397 [doi:10.14471/2017.37.0 10]
    • Griewald Y, Clemens G, Kamp J, Gladun E, Hölzel N, von Dressler H (2017) Developing land use scenarios for stakeholder participation in Russia. Land Use Policy 68: 264–276 [doi:10.1016/j.landusepol.2017.07.049]
    • Klaus VH, Schäfer D, Kleinebecker T, Fischer M, Prati D, Hölzel N (2017) Enriching plant diversity in grasslands by large-scale experimental sward disturbance and seed addition along gradients of land-use intensity. Journal of Plant Ecology 10 (4): 581–591 [doi:10.1093/jpe/rtw062]
    • Palpurina S, Wagner V, von Wehrden H, Hájek M, Hórsak M, Brinkert A, Hölzel N, Wesche K, Kamp J, Hájková P, Danihelka J, Lustyk P, Merunková K, Preislerová Z, Kočí M, Kubešová S, Cheresov M, Ermakov N, German D, Gogoleva P, Lashchynsky N, Martynenko V, Chytrý M (2017) The relationship between plant species richness and soil pH vanishes with increasing aridity across Eurasian dry grasslands. Global Ecology and Biogeography 26 (4): 425–434 [doi:10.1111/geb.12549]
    • Velbert F, Kleinebecker T, Mudrák O, Schwartze P, Hölzel N (2017) Time lags in functional response to management regimes – evidence from a 26-year field experiment in wet meadows. Journal of Vegetation Science 28 (2): 313–324 [doi:10.1111/jvs.12497]
    • Wagner V, Chytrý M, Zelený D, von Wehrden H, Brinkert A, Danihelka J, Jansen F, Hölzel N, Kamp J, Lustyk P, Merunková K, Palpurina S, Preislerová Z, Wesche K (2017) Regional differences in soil pH niche among dry grassland plants in Eurasia. Oikos 126(5): 660–670 [doi:10.1111/oik.03369]
    • Wertebach T-M, Hölzel N, Kämpf I, Yurtaev A, Tupitsin S, Kiehl K, Kamp J, Kleinebecker T (2017) Soil carbon sequestration due to post-Soviet cropland abandonment: estimates from a large-scale soil organic carbon field inventory. Global Change Biology 23 (9): 3729–3741 [doi:10.1111/gcb.13650]
    • Wolff C, Gilhaus K, Hölzel N, Schneider S (2017) Status and restoration potential of heathlands and sand grasslands in the southwest of Luxembourg. Tuexenia 37: 179–200 [doi:10.14471/2017.37.007]


    • Blüthgen N, Simons N, Jung K, Prati D, Renner S, Boch S, Fischer M, Hölzel N, Klaus VH, Kleinebecker T, Tschapka M, Weisser WW, Gossner MM (2016) Land use imperils plant and animal community stability through changes in asynchrony rather than diversity. Nature Communications 7: 10697 [doi:10.1038/ncomms10697]
    • Brinkert A, Hölzel N, Sidorova TV, Kamp J (2016) Spontaneous steppe restoration on abandoned cropland in Kazakhstan: grazing affects successional pathways. Biodiversity and Conservation 25 (12): 2543–2561 [doi:10.1007/s10531-015-1020-7]
    • Bucharova A, Durka W, Hermann J-M, Hölzel N, Michalski S, Kollmann J,  Bossdorf O (2016) Plants adapted to warmer climate do not outperform regional plants during a natural heat wave. Ecology and Evolution 6: 4160–4165 [doi:10.1002/ece3.2183]
    • Fleischer E, Khashimov I, Hölzel N, Klemm O (2016) Carbon exchange fluxes over peatlands in Western Siberia: Possible feedback between land-use change and climate change. Science of the Total Environment 545–546: 424–433 [doi:10.1016/j.scitotenv.2015.12.073]
    • Gilhaus K, Hölzel N (2016) Seasonal variations of fodder quality and availability as constraints for stocking rates in year-round grazing schemes. Agriculture, Ecosystems and Environment 234: 5–15  [doi:10.1016/j.agee.2016.03.013] - Special Issue 


    North Sea Region Climate Change Assessment - Cover
    © SpringerOpen
    • Hölzel N, Hickler T, Kutzbach L, Joosten H, van Huissteden J, Hiederer R (2016) Environmental Impacts - Terrestrial Ecosystems. In: Quante M, Colijn F (eds.) North Sea Region Climate Change Assessment. Regional Climate Studies, Part III, 341–372 [doi:10.1007/978-3-319-39745-0_11] Available under Open Access
    • Kämpf I, Hölzel N, Kiehl K (2016) Arable weed communities in the south of Western Siberia – impoverished species pools constrain diversity despite low land-use intensity. Tuexenia 36: 249–270 [doi:10.14471/2016.36.012]
    • Kämpf I, Hölzel N, Störrle M, Broll G, Kiehl K (2016) Potential of temperate agricultural soils for carbon sequestration: A meta-analysis of land-use effects. Science of The Total Environment 566–567: 428–435 [doi:10.1016/j.scitotenv.2016.05.067]
    • Kämpf I, Mathar W, Kuzmin I, Hölzel N, Kiehl K (2016) Post-Soviet Recovery of grassland vegetation on abandoned fields in the forest steppe zone of Western Siberia. Biodiversity and Conservation 25 (12): 2563–2580 [doi:10.​1007/​s10531-016-1078-x]
    • Klaus VH, Boch S, Boeddinghaus RS, Hölzel N, Kandeler E, Marhan S, Oelmann Y, Prati D, Regan KM, Schmitt B, Sorkau E, Kleinebecker T (2016) Temporal and small-scale spatial variation in grassland productivity, biomass quality and nutrient limitation. Plant Ecology 217 (7): 843–856 [doi:10.1007/s11258-016-0607-8]
    • Klaus VH*, Hölzel N, Prati D, Schmitt B, Schöning I, Schrumpf M, Solly EF, Hänsel F, Fischer M, Kleinebecker T* (2016) Plant diversity moderates drought stress in grasslands: Implications from a large real-world study on 13C natural abundances. Science of the Total Environment 566–567: 215–222 [doi:10.1016/j.scitotenv.2016.05.008] (* equal authorship)
    • Mathar W, Kämpf I, Kleinebecker T, Kuzmin I, Tolstikov A, Tupitsin S, Hölzel N (2016) Floristic diversity of meadow steppes in the Western Siberian Plain: effects of abiotic site conditions, management and landscape structure. Biodiversity & Conservation 25 (12): 2361–2379 [doi:10.1007/s10531-015-1023-4] - Special Issue
    • Rupprecht D, Gilhaus K, Hölzel N (2016) Effects of year-round grazing on the vegetation of nutrient-poor grass- and heathlands—Evidence from a large-scale survey. Agriculture, Ecosystems and Environment 234: 16–22 [doi:10.1016/j.agee.2016.02.015] - Special Issue
    • Soliveres S, Manning P, Prati D, Gossner MM, Alt F, Arndt H, Baumgartner V, Binkenstein J, Birkhofer K, Blaser S, Blüthgen N, Boch S, Böhm S, Börschig C, Buscot F, Diekötter T, Heinze J, Hölzel N, Jung K, Klaus VH, Klein A-M, Kleinebecker T, Klemmer S, Krauss J, Lange M, Morris EK, Müller J, Oelmann Y, Overmann J, Pašalić E, Renner SC, Rillig MC, Schaefer HM, Schloter M, Schmitt B, Schöning I, Schrumpf M, Sikorski J, Socher SA, Solly EF, Sonnemann I, Sorkau E, Steckel J, Steffen-Dewenter I, Stempfhuber B, Tschapka M, Türke M, Venter P, Weiner CN, Weisser WW, Werner M, Westphal C, Wilcke W, Wolters V, Wubet T, Wurst S, Fischer M, Allan E (2016) Locally rare species influence grassland ecosystem multifunctionality. Philosophical Transactions of the Royal Society B 371: 20150269 [doi:10.1098/rstb.2015.0269 ]
    • Soliveres S, van der Plas F, Manning P, Prati D, Gossner MM, Renner SC, Alt F, Arndt H, Baumgartner V, Binkenstein J, Birkhofer K, Blaser S, Blüthgen N, Boch S, Böhm S, Börschig C, Buscot F, Diekötter T, Heinze J, Hölzel N, Jung K, Klaus VH, Kleinebecker T, Klemmer S, Krauss J, Lange M, Morris EK, Müller J, Oelmann Y, Overmann J, Pašalić E, Rillig MC, Schaefer HM, Schloter M, Schmitt B, Schöning I, Schrumpf M, Sikorski J, Socher SA, Solly EF, Sonnemann I, Sorkau E, Steckel J, Steffan-Dewenter I, Stempfhuber B, Tschapka M, Türke M, Venter P, Weiner CN, Weisser WW, Werner M, Westphal C, Wilcke W, Wolters V, Wubet T, Wurst S, Fischer M, Allan E (2016) Biodiversity at multiple trophic levels is needed for ecosystem multifunctionality. Nature 536: 456–459 [doi:10.1038/nature19092]
    • Török P, Hölzel N, van Diggelen R, Tischew S (2016) Grazing in European open landscapes: How to reconcile sustainable land management and biodiversity conservation? Agriculture, Ecosystems and Environment 234: 1–4 [doi:10.1016/j.agee.2016.06.012] - Special Issue
    • Weking S, Kämpf I, Mathar W, Hölzel N (2016) Effects of land use and landscape patterns on Orthoptera communities in the Western Siberian forest steppe. Biodiversity and Conservation 25 (12): 2341–2359 [doi:10.​1007/​s10531-016-1107-9]
    • Wertebach T-M, Knorr K-H, Lordieck M, Tretiakov N, Blodau C, Hölzel N, Kleinebecker T (2016) Relationships between vegetation succession, pore water chemistry and CH₄ and CO₂ production in a transitional mire of Western Siberia (Tyumen Oblast). Wetlands 36 (5): 836–874 [doi:10.1007/s13157-016-0798-8] [free online access via Springer Nature SharedIt]


    • Allan E, Manning P, Prati D, Grassein F, Alt F, Binkenstein J, Blaser S, Blüthgen N, Hölzel N, Klaus VH, Kleinebecker T, Morris E-K, Oelmann Y, Renner SC, Rillig MC, Schäfer M, Schloter M, Schmitt B, Schöning I, Schrumpf M, Solly E, Sorkau E, Steckel J, Steffen-Dewenter I, Stempfhuber B, Trumbore S, Weiner CN, Weisser WW, Werner W, Westphal C, Wilcke W, Fischer M (2015) Land use intensification alters ecosystem multifunctionality via loss of biodiversity and changes to functional composition. Ecology Letters 18: 834–843 [doi:10.1111/ele.12469]
    • Freitag M, Mathar W, Yurtaev AA, Hölzel N (2015) Floristic composition and environmental determinants of pine forests in the hemiboreal zone of Western Siberia. Tuexenia 35: 31–51 [doi:10.14471/2015.35.017] [awarded the price for young authors by the Floristisch-soziologische Arbeitsgemeinschaft 2017]
    • Gilhaus K, Vogt V, Hölzel N (2015) Restoration of sand grasslands by topsoil removal and self-greening. Applied Vegetation Science 18 (4): 661–673 [doi:10.1111/avsc.12176]
    • Kamp J, Oppel S, Ananin AA, Durnev YA, Gashev SN, Hölzel N, Mishchenko AL, Pessa J, Smirenski SM, Strelnikov EG, Timonen S, Wolanska K, Chan S (2015) Global population collapse in a superabundant migratory bird and illegal trapping in China. Conservation Biology 29: 1684–1694 [doi:10.1111/cobi.12537]
    • Manning P, Gossner M, Bossdorf O, Allan E, Prati D, Blüthgen N, Boch S, Böhm S, Börschig C, Hölzel N, Jung K, Klaus VH, Klein A, Kleinebecker T, Krauss J, Lange M, Müller J, Pašalić E, Socher S, Tschapka M, Türke M, Weiner C, Werner M, Gockel S, Hemp A, Renner S, Wells K, Buscot F, Kalko E, Linsenmair KE, Weisser WW, Fischer M (2015) Grassland management intensification weakens the associations among the diversities of multiple plant and animal taxa. Ecology 96:1492–1501 [doi:10.1890/14-1307.1]
    • Mathar W, Kleinebecker T, Hölzel N (2015) Environmental variation as a key process of co-existence in flood-meadows. Journal of Vegetation Science 26: 480–491 [doi:10.1111/jvs.12254]


    • Allan E, Bossdorf O, Dormann CF, Prati D, Gossner M, Tscharntke T, Blüthgen N, Bellach M, Birkhofer K, Boch S, Böhm S, Börschig C, Chatzinotas A, Christ S, Daniel R, Diekoetter T, Fischer C, Friedl T, Glaser K, Hallman C, Hodac L, Hölzel N, Jung K, Klein A-M, Klaus VH, Kleinebecker T, Krauss J, Lange M, Morris EK, Müller J, Nacke H, Pašalic E, Rillig M, Rothenwöhrer C, Schall P, Scherber C, Schulze W, Socher S, Steckel J, Steffan-Dewenter I, Türke M, Weiner CN, Werner M, Westphal C, Wolters V, Wubet T, Gocke S, Gorke M, Hemp A, Renner SC, Schöning I, Pfeiffer S, König-Ries B, Buscot F, Linsenmair K-E, Schulze E-D, Weisser WW, Fischer M (2014) Inter-annual variation in land-use intensity enhances grassland multidiversity. Proceedings of the National Academy of Sciences of the United States of America 111 (1):  308–313 [doi:10.1073/pnas.1312213111]
    • Fujita Y, Olde Venterink H, van Bodegom PM, Douma JC, Heil GW, Hölzel N, Jabłońska E, Kotowski W, Okruszko T, Pawlikowski P, de Ruiter PC, Wassen MJ (2014) Low investment in sexual reproduction threatens plants adapted to phosphorus limitation. Nature 505: 82–86 [doi:10.1038/nature12733]
    • Gilhaus K, Stelzner F, Hölzel N (2014) Cattle foraging habits shape vegetation patterns of alluvial year-round grazing systems. Plant Ecology 215 (2): 169–179 [doi:10.1007/s11258-013-0287-6]
    • Kleinebecker T, Hölzel N, Prati D, Schmitt B, Fischer M, Klaus VH (2014) Evidence from the real world: 15N natural abundances reveal enhanced nitrogen use at high plant diversity in Central European grasslands. Journal of Ecology 102 2: 456–465 [doi:10.1111/1365-2745.12202]
    • Walker LR, Hölzel N, Marrs R, del Moral R, Prach K (2014) Optimization of intervention levels in ecological restoration. Applied Vegetation Science 17 (2): 187–192 [doi:10.1111/avsc.12082]
    • Wertebach T-M, Hölzel N, Kleinebecker T (2014) Birch encroachment affects the base cation chemistry in a restored bog. Ecohydrology 7 (4): 1163–1171 [doi:10.1002/eco.1447]


    • Jedrzejek B, Drees B, Daniëls FJA, Hölzel N (2013) Vegetation pattern of mountains in West Greenland – a baseline for long-term surveillance of global warming impacts. Plant Ecology & Diversity 6(3-4): 405–422 [doi:10.1080/17550874.2013.802049]
    • Klaus VH, Hölzel N, Boch S, Müller J, Socher SA, Prati D, Fischer M, Kleinebecker T (2013) Direct and indirect associations between plant species richness and productivity in grasslands: regional differences preclude simple generalization of productivity-biodiversity relationships. Preslia 85: 97–112 [link open access pdf]
    • Klaus VH, Hölzel N, Prati D, Schmitt B, Schöning I, Schrumpf M, Fischer M, Kleinebecker T (2013) Organic vs. conventional grassland management: Do 15N and 13C isotopic signatures of hay and soil samples differ? PLoS ONE 8(10): e78134 [doi:10.1371/journal.pone.0078134]
    • Klaus VH, Kleinebecker T, Prati D, Fischer M, Alt F, Boch S, Gockel S, Gossner M, Hemp A, Lange M, Müller J, Oelmann Y, Pašalić E, Renner SC, Socher SA, Türke M, Weisser WW, Hölzel N (2013) Does organic grassland farming benefit plant and arthropod diversity at the expense of yield and soil fertility? Agriculture, Ecosystems and Environment 177:1–9 [doi:10.1016/j.agee.2013.05.019]
    • Kleinebecker T, Poelen MDM, Smolders AJP, Lamers LPM, Hölzel N (2013) Fast and Inexpensive Detection of Total and Extractable Element Concentrations in Aquatic Sediments Using Near-Infrared Reflectance Spectroscopy (NIRS). PLoS ONE 8 (7): e70517 [doi:10.1371/journal.pone.0070517]
    • Neuenkamp L, Metsoja JA, Zobel M, Hölzel N (2013) Impact of management on biodiversity-biomass relations in Estonian flooded meadows. Plant Ecology 214 (6): 845–856 [doi:10.1007/s11258-013-0213-y]
    • Teuber LM, Hölzel N, Fraser LH (2013) Livestock grazing in intermountain depressional wetlands - Effects on plant strategies, soil characteristics and biomass. Agriculture, Ecosystems and Environment 175: 21–28 [doi:10.1016/j.agee.2013.04.017]


    • Berndt A, Hölzel N (2012) Energy crops as a new bird habitat: Utilization of oilseed rape fields by the rare Bluethroat (Luscinia svecica). Biodiversity and Conservation 21: 527–541 [doi:10.1007/s10531-011-0199-5]
    • Blüthgen N, Dormann C, Prati D, Klaus VH, Kleinebecker T, Hölzel N, Alt F, Boch S, Gockel S, Hemp A,  Müller J, Nieschulze J, Renner SC, Schöning I, Schumacher U, Socher SA, Wells K, Birkhofer K, Buscot F,  Oelmann Y, Rothenwöhrer C, Scherber C, Tscharntke T,  Weiner C, Fischer M, Kalko EKV, Linsenmair KE, Schulze E-D, Weisser WW (2012) A quantitative index of land-use intensity in grasslands: integrating mowing, grazing and fertilization. Basic and Applied Ecology 13: 207–220 [doi:10.1016/j.baae.2012.04.001]
    • Applied Vegetation Science 2012 - Special Issue: Vegetation Restoration, Edited by Norbert Hölzel, Elise Buisson & Thierry Dutoit - CoverHölzel N, Buisson E, Dutoit T (2012): Species introduction - a major topic in vegetation restoration. Applied Vegetation Science 15 (2): 161–165 [doi:10.1111/j.1645-109X.2012.01189.x]
      AVS Special Issue: Vegetation Restoration, Edited by Norbert Hölzel, Elise Buisson & Thierry Dutoit
    • Jedrzejek B, Drees B, Daniëls FJA, Hölzel N (2012) Vegetation discontinuities and altitudinal indicator species in mountains of West Greenland: finding the best positions and traits to observe the impact of climate warming in the Arctic. Applied Vegetation Science 15: 432–448 [doi:10.1111/j.1654-109X.2012.01186.x]
    • Kamp J, Siderova T, Salemgaarev A, Urazaliev R, Donald PF, Hölzel N (2012) Niche separation of larks (Alaudidae) and agricultural change on the drylands of the former Soviet Union. Agriculture, Ecosystems and Environment 155: 41–49 [doi: 10.1016/j.agee.2012.03.023]
    • Klaus VH, Kleinebecker T, Boch S, Müller J, Socher AS, Prati D, Fischer M, Hölzel N (2012) NIRS meets Ellenberg's indicator values: Prediction of moisture and nitrogen values of agricultural grassland vegetation by means of near-infrared spectral characteristics. Ecological Indicators 14: 82–86 [doi:10.1016/j.ecolind.2011.07.016]
    • Müller J, Klaus VH, Kleinebecker T, Prati D, Hölzel N, Fischer M (2012) Impact of Land-Use Intensity and Productivity on Bryophyte Diversity in Agricultural Grasslands. PLoS ONE 7 (12): e51520 [doi:10.1371/journal.pone.0051520]
    • Socher S, Prati D, Müller J, Klaus V, Hölzel N, Fischer M (2012) Direct and productivity-mediated indirect effects of fertilization, mowing and grazing intensities on grassland plant species richness. Journal of Ecology 100: 1391–1399 [doi:10.1111/j.1365-2745.2012.02020.x]


    • Kamp J, Urazaliev R, Donald PF, Hölzel, N (2011) Post-Soviet agricultural change predicts future declines after recent recovery in Eurasian steppe bird populations. Biological Conservation 144: 2607–2614 [doi:10.1016/j.biocon.2011.07.010]
    • Klaus VH, Kleinebecker T, Hölzel N, Blüthgen N, Boch S, Müller J, Socher SA, Prati D, Fischer M (2011) Nutrient concentrations and fibre contents of plant community biomass reflect species richness patterns along a broad range of land-use intensities among agricultural grasslands. Perspectives in Plant Ecology, Evolution and Systematics 13: 287–295 [doi:10.1016/j.ppees.2011.07.001]
    • Klaus VH, Sintermann J, Kleinebecker T, Hölzel N (2011) Sedimentation-induced eutrophication in large river floodplains - an obstacle to restoration? Biological Conservation 144: 451–458 [doi:10.1016/j.biocon.2010.09.031]
    • Kleinebecker T, Klaus VH, Hölzel N (2011) Reducing sample quantity and maintaining high prediction accuracy of quality parameters in grassland biomass with near-infrared reflectance spectroscopy (NIRS). Journal of Near Infrared Spectroscopy 19: 495–505 [doi:10.1255/jnirs.957]
    • Kleinebecker T, Weber H, Hölzel N (2011) Effects of grazing on seasonal variation of aboveground biomass quality in calcareous grasslands. Plant Ecology (2011) 212: 1563–1576 [doi:10.1007/s11258-011-9931-1]


    • Kiehl K, Kirmer A, Donath T, Rasran L, Hölzel N (2010) Species introduction in restoration projects - method evaluation for the establishment of semi-natural grasslands. Basic and Applied Ecology 11: 285–299 [doi:10.1016/j.baae.2009.12.004]
    • Kleinebecker T, Hölzel N, Vogel A (2010) Patterns and gradients of diversity in South Patagonian ombrotrophic peat bogs. Austral Ecology 35: 1–12 [doi:10.1111/j.1442-9993.2009.02003.x]
    • Schmidt SR, Kleinebecker T, Vogel A, Hölzel N (2010) Interspecific and geographical differences of plant tissue nutrient concentrations along an environmental gradient in Southern Patagonia, Chile. Aquatic Botany 92: 149–156 [doi:10.1016/j.aquabot.2009.11.002]


    • Kleinebecker T, Schmidt SR, Fritz C, Smolders AJP, Hölzel N (2009) Prediction of δ13C and δ15N in plant tissues with near-infrared reflectance spectroscopy. New Phytologist 184: 732–739 [doi:10.1111/j.1469-8137.2009.02995.x
    • Oelmann Y, Broll G, Hölzel N, Kleinebecker T, Vogel A, Schwartze P (2009) Nutrient impoverishment and limitation of productivity after 20 years of conservation management in wet grasslands of north-western Germany. Biological Conservation 142: 2941–2948 [abstract ScienceDirect]
    • Poptcheva K, Schwartze P, Vogel A, Kleinebecker T, Hölzel N (2009) Changes in wet meadow vegetation after 20 years of different management in a field experiment (North-West Germany). Agriculture, Ecosystems and Environment 134: 108–114 [abstract ScienceDirect]


    • JVS Cover Rio RubensKleinebecker T, Hölzel N, Vogel A (2008) South Patagonian ombrotrophic bog vegetation reflects biogeochemical gradients at the landscape level. Journal of Vegetation Science 19: 151–160 [pdf 191 KB]


    • Donath TW, Bissels S, Hölzel N, Otte A (2007) Large scale application of diaspore transfer with plant material in restoration practice - Impact of seed and microsite limitation. Biological Conservation 138: 224–234 [abstract ScienceDirect]
    • Kleinebecker T, Hölzel N, Vogel A (2007) Gradients of continentality and moisture in South Patagonian ombrotrophic peatland vegetation. Folia Geobotanica 42: 363–382 [pdf 385 KB]


    • Bissels S, Donath TW, Hölzel N, Otte A (2006) Effects of different mowing regimes on seedling recruitment in alluvial meadows. Basic and Applied Ecology 7: 433–442 [abstract ScienceDirect]
    • Donath TW, Hölzel N, Otte A (2006) Influence of competition by sown grass, disturbance and litter on recruitment of rare flood-meadow species. Biological Conservation 130: 315–323 [abstract ScienceDirect]
    • Eckstein RL, Hölzel N, Danihelka J (2006) Biological Flora of Central Europe: Viola elatior, V. pumila, and V. stagnina. Perspectives in Plant Ecology, Evolution and Systematics 8: 45–66 [abstract ScienceDirect]


    • Bissels S, Donath TW, Hölzel N, Otte A (2005) Ephemeral wetland vegetation of irregularly flooded arable fields: The importance of persistent soil seed banks. Phytocoenologia 35/2-3: 469–488 [abstract IngentaConnect]
    • Hölzel N (2005) Seedling recruitment in flood-meadow species: The effects of gaps, litter and vegetation matrix. Applied Vegetation Science 8: 115–124 [pdf 717 KB]


    • Bissels S, Hölzel N, Otte A (2004) Population structure of the threatened long-lived perennial Serratula tinctoria in relation to vegetation and management. Applied Vegetation Science 7: 267–274 [pdf 367 KB]
    • Bissels S, Hölzel N, Donath TW, Otte A (2004) Evaluation of restoration success in alluvial grasslands under contrasting flooding regimes. Biological Conservation 18: 641–650 [abstract ScienceDirect]
    • Donath TW, Hölzel N, Bissels S, Otte A (2004) Perspectives for incorporating biomass from non-intensively managed temperate flood meadows into farming systems. Agriculture, Ecosystems and Environment 104: 439–451 [abstract ScienceDirect]
    • Eckstein RL, Danihelka J, Hölzel N, Otte A (2004) The effects of environmental variation and management on population structure in three river-corridor violets. Acta Oecologica 25: 83–91 [abstract ScienceDirect]
    • Hölzel N, Otte A (2004a) Assessing soil seed bank persistence in flood-meadows: The search for reliable traits. Journal of Vegetation Science 15: 93–100 [pdf 127 KB appendix 54 KB]
    • Hölzel N, Otte A (2004b) Ecological significance of seed germination characteristics in flood-meadow species. Flora 199: 12–24 [doi:10.1078/0367-2530-00132] [pdf 121 KB abstract ScienceDirect]
    • Hölzel N, Otte A (2004c) Inter-annual variation in the soil seed bank of flood-meadows over two years with different flooding patterns. Plant Ecology 174: 279–291 [doi: 10.1023/B:VEGE.0000049108.04955.e2] [pdf 209 KB]


    • Donath TW, Hölzel N, Otte A (2003) The impact of site conditions and seed dispersal on restoration success in alluvial meadows. Applied Vegetation Science 6: 13–22 [doi: 10.1111/j.1654-109X.2003.tb00560.x] [pdf 262 KB appendix 10 KB]
    • Hölzel N (2003) Re-assessing the ecology of rare flood-meadow violets (Viola elatior, V. pumila, V. persicifolia) with large phytosociological data sets. Folia Geobotanica 38: 281–298  [doi: 10.1007/BF02803200] [pdf 178 KB]
    • Hölzel N, Otte A (2003) Restoration of a species-rich flood-meadow by topsoil removal and diaspore transfer with plant material. Applied Vegetation Science 6: 131–140 [doi: 10.1111/j.1654-109X.2003.tb00573.x] [awarded ‘paper of the year’ by the chief editors of Applied Vegetation Science] [pdf 207 KB appendix 15 KB Editor's Award 175 KB]


    • Hölzel N, Haub C, Ingelfinger MP, Otte A, Pilipenko VN (2002) The return of the steppe – large-scale restoration of degraded land in southern Russia during the post-Soviet era. Journal for Nature Conservation 10: 75–86 [doi:10.1078/1617-1381-00009]


    • Hölzel N, Otte A (2001) The impact of flooding regime on the soil seed bank of flood-meadows. Journal of Vegetation Science 12: 209–218 [doi:10.2307/3236605] [pdf 63 KB appendix 15 KB]

    books and book chapter

    • Zerbe Wiegleb 2009 Renaturierung von Ökosystemen in MitteleuropaHölzel N, mit Beiträgen von Rebele F, Rosenthal G, Eichberg C (2009) Ökologische Grundlagen und limitierende Faktoren der Renaturierung. In: Zerbe S, Wiegleb G (Hrsg) Renaturierung von Ökosystemen in Mitteleuropa. Spektrum Akademischer Verlag, Heidelberg: 23–53 [read online at springerLink]
    • Rosenthal G & Hölzel N (2009) Renaturierung von Feuchtgrünland, Auengrünland und mesophilem Grünland. In: Zerbe S & Wiegleb G (Hrsg) Renaturierung von Ökosystemen in Mitteleuropa. Spektrum Akademischer Verlag, Heidelberg: 283–316 [read online at springerLink]
    • Härdtle W, Ewald J, Hölzel N (2004) Wälder des Tieflandes und der Mittelgebirge.  Ökosysteme Mitteleuropas aus geobotanischer Sicht (Hrsg. Pott R), Eugen Ulmer: 252 S, Stuttgart

    other publications

    • Hölzel N, Klaus VH (2017) Zur Artenvielfalt im Grünland. Was sie beeinflusst, wozu wir sie brauchen und wie wir sie unterstützen können. Natur in NRW 2/2017: 35–39 [download]
    • Boch S, Allan E, Birkhofer K, Blüthgen N, Christ-Breulmann S, Diekoetter T, Dormann CF, Gossner MM, Hallmann C, Hemp A,  Hölzel N, Jung K, Klaus VH,  Klein A-M, Kleinebecker T, Lange M, Müller J, Nacke H, Prati D, Renner SC, Rothenwöhrer C, Schall P, Schulze E-D, Socher SA, Tscharntke T, Türke M, Weiner CN, Weisser WW, Westphal C, Wolters V, Wubet T, Fischer M (2016): Extensive und jährlich wechselnde Nutzungsintensität fördert den Artenreichtum im Grünland. ANLiegen Natur 38 (1): online, 10pp [download]
    • Fleischer K, Hölzel N, Eds. (2012) Jahrestagung der Floristisch-soziologischen Arbeitsgemeinschaft (FlorSoz) in Münster 2012 mit Exkursionen in Westfalen und den benachbarten Niederlanden. Tuexenia, Beiheft Nr. 5
    • Fartmann T, Behrens M, Möllenbeck V, Hölzel N (2012) Potential effects of climate change on the biodiversity in North Rhine-Westphalia. In: Ellwanger G, Ssymank A & Paulsch C (Hrsg.) Natura 2000 and Climate Change – a Challenge. Naturschutz und Biologische Vielfalt 118: 63–72
    • Möllenbeck V, Behrens M, Fartmann T, Hölzel N, Hübner T, Kiel E-F (2011) Auswirkungen des Klimawandels auf die Biologische Vielfalt. Empfindlichkeitsanalysen für ausgewählte Tier- und Pflanzenarten und Lebensräume in Nordrhein-Westfalen. Natur in NRW 4/11: 15–19 [Link zum pdf]
    • Hölzel N (2011) Artenanreicherung durch Mahdgutübertragung - Möglichkeiten und Grenzen der Mahdgutübertragung. Natur in NRW 2/11: 22–24 [Link pdf]
    • Sieg B, Kleinebecker T, Hölzel N (2010) Evaluation of large scale bog restoration in North-Western Germany. Lessons from 30 years of practice. Extended abstract 7th European Conference on Ecological Restoration, Avignon, France. SER Europe Knowledge Base (www.ser.org/europe), 4 pp
    • Schmitt B, Fartmann T, Hölzel N (2010) Vergesellschaftung und Ökologie der Sumpf-Siegwurz (Gladiolus palustris) in Südbayern. Tuexenia 30: 105–127 [Download www.tuexenia.de] [awarded the price for young authors by the Floristisch-soziologische Arbeitsgemeinschaft 2010]
    • Graef F, Bilo M, Weddeling K, Hölzel N (eds.) (2009) Einsatz von Fernerkundung im Rahmen des FFH-Monitorings in Deutschland. BfN-Skripten 249: 1–130 [Download www.bfn.de]
    • Kleinebecker T, Vogel A & Hölzel N (2008) Die Vegetation ombrotropher Moore Südpatagoniens. Abhandlungen aus dem westfälischen Museum für Naturkunde 70: 439–452 [pdf 1,1 MB]
    • Donath TW, Bissels S, Handke K, Harnisch M, Hölzel N, Otte A (2006) E + E-Vorhaben "Stromtalwiesen" - Renaturierung von Stromtalwiesen am hessischen Oberrhein durch Mahdgutübertragung. Natur und Landschaft 11: 529–535
    • Hölzel N, Bissels S, Donath TW, Handke K, Harnisch M, Otte A (2006) Renaturierung von Stromtalwiesen am hessischen Oberrhein - Ergebnisse aus dem E + E-Vorhaben 89211-9/00 des Bundesamtes für Naturschutz. Naturschutz und Biologische Vielfalt 31: 1–263
    • Burkart M, Dierschke H, Hölzel N, Nowak B, Fartmann T (2004) Molinio-Arrhenatheretea (E1) - Kulturgrasland und verwandte Vegetationstypen; Teil 2: Molinietalia - Futter- und Streuwiesen feucht-nasser Standorte. Synopsis der Pflanzengesellschaften Deutschlands, Heft 9: 103 S, Göttingen
    • Hölzel N, Schwabe-Kratochwil A, Rosenthal G (2004) Symposium 10: Constraints and perspectives of grassland restoration. Verhandlungen der Gesellschaft für Ökologie 34: 148–150
    • Hölzel N & Harnisch M (2002) Renaturierung von Stromtalwiesen. Garten und Landschaft 112/8: 36–38
    • Hölzel N, Donath TW, Bissels S & Otte A (2002) Auengrünlandrenaturierung am hessischen Oberrhein – Defizite und Erfolge nach 15 Jahren Laufzeit. Schriftenreihe für Vegetationskunde 36: 131-137
    • Pfadenhauer J, Ewald J, Hölzel N (2001) Northern limestone prealps: Brauneck and Benediktenwand Mts. In: Pfadenhauer J, Albrecht H & Erschbamer B (eds.) Post-symposium excursion, 44th IAVS Symposium 29 July – 4 August 2001 Freising-Weihenstephan, Germany, S. 46–65. Department of Ecology TUM, Freising
    • Hölzel N (1999a) Flora und Vegetation der Auenwiesen im NSG „Lampertheimer Altrhein“ – eine aktuelle Zustandsanalyse mit Hinweisen zur zukünftigen Pflege und Entwicklung. Jahrbuch Naturschutz in Hessen 4: 24–42
    • Hölzel N (1999b) Geobotanische Dauerbeobachtung als Grundlage für die Effizienzkontrolle von Naturschutzmaßnahmen in Streuobstwiesen. Naturschutz und Landschaftsplanung 31/5: 147–154
    • Hölzel N (1998) Pflege und Entwicklung mahdgeprägter, orchideenreicher Trespen-Halbtrockenrasen an der Hessischen Bergstraße durch eine Naturschutzgruppe. – Jahrbuch Naturschutz in Hessen 3: 117–120
    • Otte A, Hölzel N (1998) „Biotop des Jahres“ – Die Obstwiese – Zustandstandsbericht über die „Maintaler Streuobstwiesen“. Lebensraum – Zeitschrift für Naturschutz in der Kulturlandschaft 10/2: 4–5
    • Russanow GM, Hölzel N, Schleuning S (1998) Siberian White Crane in Volga-delta. Dutch Birding 20/3: 101–106
    • Hölzel N (1997) Wolga-Delta: Paradies mit ökologischen Problemen. - GLOBUS Begleitheft zur ARD-Fernsehreihe aus Natur und Umwelt, 16. Jahrgang, Doppelheft 4-5/97: 76–83, Stuttgart
    • Hölzel N (1996a) Schneeheide-Kiefernwälder in den mittleren Nördlichen Kalkalpen. Laufener Forschungsberichte 3: 192 S, Laufen
    • Hölzel N (1996b) Erico-Pinetea (H6) - Alpisch-Dinarische Karbonat-Trocken-Kiefernwälder. Synopsis der Pflanzengesellschaften Deutschlands Heft 1: 11–49
    • Hölzel N (1996c) Zur floristischen Struktur, Ökologie und Dynamik alpischer Karbonat-Trockenkiefernwälder der Klasse Erico-Pinetea. Berichte der Reinhold-Tüxen Gesellschaft 8: 79–98
    • Hölzel N (1996d) Zur Verbreitung und Ökologie von Karbonat-Trockenkiefernwäldern (Erico-Pinetea) in den Bayerischen und Nordtiroler Kalkalpen. Berichte der Naturwissenschaftlichen Gesellschaft Bayreuth XXIII: 492-495
    • Hölzel N, Fischer A (1996) Buntreitgraskiefernwälder im Werdenfelser Land. In: Pfadenhauer J & Albrecht H (Hrsg.) Floristisch-soziologische Arbeitsgemeinschaft: Exkursionsführer zur 46. Jahrestagung der Floristisch-soziologischen Arbeitsgemeinschaft vom 12. – 15. Juli in Freising-Weihenstephan: 245–261, Eching (IHW)
    • Hölzel N, Russanow G, Schleuning S (1996) Wolga-Delta - Naturoase zwischen Meer und Halbwüste. Naturerbe Verlag Jürgen Resch, 159 S, Überlingen
    • Otte A, Seibert P, Hölzel N (1996a) Die Donauniederung zwischen Neuburg und Ingolstadt: Sequenz der Auenvegetation. In: Pfadenhauer J & Albrecht H (Hrsg) Floristisch-soziologische Arbeitsgemeinschaft: Exkursionsführer zur 46. Jahrestagung der Floristisch-soziologischen Arbeitsgemeinschaft vom 12. – 15. Juli in Freising-Weihenstephan: 179–191, Eching (IHW)
    • Otte A, Seibert P, Hölzel N (1996b) Vegetationsabfolge in voralpinen Flußlandschaften und ihre anthropogene Veränderung. In: Pfadenhauer J & Albrecht H (Hrsg) Floristisch-soziologische Arbeitsgemeinschaft: Exkursionsführer zur 46. Jahrestagung der Floristisch-soziologischen Arbeitsgemeinschaft vom 12. – 15. Juli in Freising- Weihenstephan: 179–191, Eching (IHW)
    • Schleuning M, M & S, Hölzel N (1997) Birding the limits of the Western Palaearctic - the Volga Delta. Birding World 10/8: 312-316, Holt, Norfolk
    • Hölzel N (1995) Dynamik von nordalpinen Trockenkiefernwäldern - Konsequenzen für Naturschutz und Landschaftspflege. Laufener Seminarbeiträge 3/95: 81–91
    • Hölzel N (1994) Standortverhältnisse, Flora und Vegetation. In: Kornprobst M: Lebensraumtyp Streuobst. - Landschaftspflegekonzept Bayern, Band II.5. - Hrsg.: Bayerisches Staatsministerium für Landesentwicklung und Umweltfragen (StMLU) und Bayerische Akademie für Naturschutz und Landschaftspflege (ANL) 21–22, 29–37, München
    • Strobel Ch, Hölzel N (1994) Lebensraumtyp Feuchtwiesen. Landschaftspflegekonzept Bayern, Band II.6. - Hrsg.: Bayerisches Staatsministerium für Landesentwicklung und Umweltfragen (StMLU) und Bayerische Akademie für Naturschutz und Landschaftspflege: 204 S, München